Ecological dynamics

The information in this ecological site description (ESD), including the state-and-transition model (STM), was developed using archeological and historical data, professional experience, and scientific studies. The information is representative of a complex set of plant communities. Not all scenarios or plants are included. Key indicator plants, animals, and ecological processes are described to inform land management decisions.

Introduction – In southeastern Texas and southwestern Louisiana the transition from coastal grasslands to the large expanse of coniferous forest has been deemed the “Flatwoods”. As the name suggests, the region is relatively flat and, with many transitional areas, highly diverse in flora and fauna. Historically, the area was covered by pines with mixed hardwoods, sparse shrubs, and a diverse understory of grasses and forbs. Fire and drainage patterns play a significant role in shaping the plant communities and their development. Fire suppression, drainage alterations, and land conversion have reduced the amount of historical communities in existence today.

Background – Prior to settlement by the Europeans, the reference state for the Poorly Drained Clayey Bottomlands were Water Oak/Overcup Oak Forests. Remnants of this presumed historic plant community still exist where natural conditions are intact. Evidence of the reference state is found in accounts of early historic explorers to the area, historic forest and biological survey teams, as well as recent ecological studies in the last 30 years. The age of this community varies and supports a diverse flora.

Settlement Management – As human settlement increased throughout the area, so did the increase in logging and grazing by domestic livestock. The logging became so extensive that by the 1930’s most of the region had been cut-over. Replanting trees to historic communities was not common and early foresters began planting loblolly pine (Pinus taeda) for its quick growth. As more people colonized they began suppressing fire, which allowed dense thickets of shrubs to replace the herbaceous understory. While this is typical for most of the region, unless the drainage patterns were altered in the swamp areas, they have retained their natural species makeup and loblolly is not often seen in the swamps.

Current Management and State – Today much of the historic forest is gone, replaced by pine plantations, crops, and pastures. The areas that were not converted have been fire-suppressed so long that loblolly pine and fire intolerant hardwoods populate the overstory structure. Currently, federally-managed properties are the best place to view the remnant sites (National Park Service, U.S. Fish and Wildlife Service, etc.). Some private individuals have begun restoring communities through selective tree planting and retention of communities that remain. Other restoration efforts include mimicking natural-disturbance regimes through gap-phase regeneration on plantation sites.

Fire Regimes – Fire was a natural and important disturbance throughout the region. Fire occurred naturally from lightning strikes, by Native Americans for game movement, and eventually early European settlers. Fires throughout the Flatwoods occurred at two different times. Early in the year, they would occur during late winter and early spring, removing senescent vegetation, recycling nutrients and minerals, and spurring new plant growth. Late summer and early fall fires occurred as well, but with a different community effect. Summer fires burned hotter and with more intensity, greatly suppressing the shrub canopy layer. The summer fires also shifted the ecological site transitional state by decreasing grass densities and increasing forb densities. The topography, fuel loads, and other conditions caused patchy burns throughout the region resulting in mosaic patterns of plant communities and a heterogeneous landscape. Fire naturally would have occurred very infrequently (once every 25 to 50 years) and only when conditions were extremely droughty.

Disturbance Regimes – Extreme weather events occur occasionally throughout the region. Tornados uproot trees and open canopies in the spring months. In the late summer and early fall, hurricanes or tropical depressions can make landfall, dumping excessive amounts of rain and toppling trees with high winds. Another cause of large canopy openings is the effects of the southern pine beetle (Dendroctonus frontalis). Starting in the late 1950’s, beetle outbreaks have occurred every 6 to 9 years (although a major attack has not occurred in some time); usually when the trees are stressed due to multiple environmental factors. Flooding is the disturbance that affects the bottomland sites most often. The amount and timing of flooding events shapes the plant communities by allowing only species adapted to the hydrologic conditions to thrive.