Natural Resources
Conservation Service
Ecological site F144BY504ME
Enriched Loamy Cove
Last updated: 9/27/2024
Accessed: 07/15/2026
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Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.
MLRA notes
Major Land Resource Area (MLRA): 144B–New England and Eastern New York Upland, Northern Part
This major land resource area (MLRA) is in Maine (56 percent), New Hampshire (22 percent), Vermont (14 percent), Massachusetts (6 percent), Connecticut (1 percent), and New York (1 percent). It makes up about 22,728 square miles (58,864 square kilometers). The MLRA consists of a relatively young landscape shaped by the Laurentide Ice Sheet, which covered the region from 35,000 to 10,000 years ago. Rolling hills of dense basal till converge on ridges of shallow bedrock that were scoured by glacial ice. River valleys that were flooded by melting glacial water or seawater house large expanses of glacial outwash and stratified drift in inland areas and, to a lesser extent, glaciomarine and glaciolacustrine sediment deposits in coastal areas. Organic bogs, ablation till, and alluvial flood plains make up the remaining portions of the MLRA.
The soils in this region are dominantly Entisols, Spodosols, and Inceptisols. They commonly have a fragipan. The dominant suborders are Ochrepts, Orthods, Aquepts, Fluvents, and Saprists. The soils in the region dominantly have a frigid soil temperature regime with some cryic areas at higher elevation, a udic soil moisture regime, and mixed mineralogy. Most of the land is forested, and 98 percent is privately owned. Significant amounts of forest products are produced including lumber, pulpwood, Christmas trees, and maple syrup. Principal agricultural crops include forage and grains for dairy cattle, potatoes, apples, and blueberries. Wildlife habitat and recreation are important land uses. Stoniness, steep slopes, and poor drainage limit the use of many of the soils.Classification relationships
NRCS:
Land Resource Region: R—Northeastern Forage and Forest Region
MLRA: 144B—New England and Eastern New York Upland, Northern PartEcological site concept
This site occurs in well-drained coves, drainageways, and other protected areas where nutrients accumulate. These are typically small inclusions within larger hardwood stands and are often not mapped as major soil components. However, the soils and vegetation are distinctive, with a thick dark-colored mineral horizon at the surface indicating high organic matter inputs. White ash, sugar maple, and yellow birch are often abundant on this site, with other rich site indicators such as royal fern and blue cohosh.
Associated sites
F144BY502ME Loamy Till Toeslope
The somewhat poorly- and poorly-drained Loamy Till Toeslope site occurs in similar landscape positions as the Enriched Loamy Cove site, but tends not to accumulate quite as many nutrients and water.
F144BY501ME Loamy Slope (Northern Hardwoods)
The Loamy Slope site typically occurs upslope of the Enriched Loamy Cove site.
Similar sites
F144BY506ME Semi-rich Till Slope
The Semi-rich Till Slope site is very similar to the Enriched Loamy Cove site, but it derives its nutrients primarily from calcareous parent material and does not accumulate nutrients due to its more exposed landscape position. Therefore it tends to be less rich than the Enriched Loamy Cove.
F144BY507ME Semi-rich Till Toeslope
The Semi-rich Till Toeslope is somewhat poorly- and poorly-drained, supporting wetter species than the Enriched Loamy Cove site. Otherwise, these two sites may be very similar.
Table 1. Dominant plant species
Tree (1) Acer saccharum
(2) Betula papyriferaShrub Not specified
Herbaceous Not specified
Physiographic features
This site occurs in small concave areas that are highly protected, such that nutrients accumulate and enrich the site. There are often small ephemeral streams and drainageways through these sites, though they also occur at the base of hills, cliffs, or other areas receiving nutrient inputs from higher in the watershed. The site occurs from sea level to 2500 feet, mostly on gentle slopes.
Table 2. Representative physiographic features
Landforms (1) Upland > Hill
Runoff class Medium to high Flooding frequency None Ponding frequency None Elevation 0 – 2500 ft Slope 0 – 35 % Water table depth 0 – 72 in Aspect Aspect is not a significant factor Climatic features
The climate is humid and temperate and is characterized by warm summers and cold winters. Precipitation generally is evenly distributed throughout the year. Near the coast, it is slightly lower in summer. In inland areas, it is slightly higher in spring and fall. Rainfall occurs during high-intensity, convective thunderstorms in summer. In winter, most of the precipitation occurs as moderate-intensity storms (northeasters) that produce large amounts of rain or snow. Heavy snowfalls commonly occur late in winter. Temperatures and the length of the freeze-free period increase from north to south and closer to the coast.
This major land resource area (MLRA) covers four states and may have substantial climate variability among locations: Maine (56 percent), New Hampshire (22 percent), Vermont (14 percent), Massachusetts (6 percent), Connecticut (1 percent), and New York (1 percent).Table 3 Representative climatic features
Frost-free period (characteristic range) 120-140 days Freeze-free period (characteristic range) 140-170 days Precipitation total (characteristic range) 40-50 in Frost-free period (actual range) 100-150 days Freeze-free period (actual range) 130-180 days Precipitation total (actual range) 40-50 in Frost-free period (average) 130 days Freeze-free period (average) 160 days Precipitation total (average) 50 in Characteristic rangeActual rangeBarLineFigure 1. Monthly precipitation range
Characteristic rangeActual rangeBarLineFigure 2. Monthly minimum temperature range
Characteristic rangeActual rangeBarLineFigure 3. Monthly maximum temperature range
BarLineFigure 4. Monthly average minimum and maximum temperature
Figure 5. Annual precipitation pattern
Figure 6 Annual average temperature pattern
Climate stations used
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(1) BELFAST [USC00170480], Belfast, ME
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(2) ACADIA NP [USC00170100], Bar Harbor, ME
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(3) CORINNA [USC00171628], Corinna, ME
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(4) DOVER-FOXCROFT WWTP [USC00171975], Dover Foxcroft, ME
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(5) FARMINGTON [USC00172765], Farmington, ME
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(6) GARDINER [USC00173046], Gardiner, ME
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(7) JONESBORO [USC00174183], Addison, ME
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(8) LEWISTON [USC00174566], Auburn, ME
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(9) MADISON [USC00174927], Anson, ME
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(10) NEWCASTLE [USC00175675], Newcastle, ME
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(11) ORONO [USC00176430], Old Town, ME
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(12) WATERVILLE TRTMT PLT [USC00179151], Waterville, ME
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(13) WEST ROCKPORT 1 NNW [USC00179593], Rockport, ME
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(14) AUGUSTA STATE AP [USW00014605], Augusta, ME
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(15) BANGOR INTL AP [USW00014606], Bangor, ME
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(16) PORTLAND INTL JETPORT [USW00014764], Portland, ME
">Influencing water features
This site is an enriched upland that may experience occasional pulses of overland flow during the wettest periods. Although it is often associated with very small ephemeral streams and drainages, this site does not function as a riparian or wetland area.
Soil features
The soils of this site are characterized by a thick, dark surface horizon high in organic matter. Soil pH ranges from 5.1 to 7.3, with high organic matter. These soils are moderately well- to well-drained with loamy textures and few rock fragments throughout. These soils are often not mapped because of their small size, so existing data is limited. However, these small patches are common and show consistent soil-vegetation patterns throughout the MLRA.
The representative soil is Herkimer.Table 4. Representative soil features
Parent material (1) Till – igneous, metamorphic and sedimentary rock
(2) Saprolite
Surface texture (1) Silt loam
Drainage class Well drained Permeability class Very slow Soil depth 40 – 0 in Surface fragment cover <=3" 0 – 2 % Surface fragment cover >3" 1 – 7 % Available water capacity
(4-6in)Not specified Soil reaction (1:1 water)
(3.5-7.3in)Not specified Subsurface fragment volume <=3"
(0-10in)Not specified Subsurface fragment volume >3"
(0-7in)Not specified Ecological dynamics
[Caveat: The vegetation information contained in this section and is only provisional, based on concepts, and future projects support validation through field work. *] The vegetation groupings described in this section are based on the terrestrial ecological system classification and vegetation associations developed by NatureServe (Comer et al., 2003) and localized associations provided by the New York Natural Heritage Program (Edinger et al., 2014), Maine Natural Areas Program (Gawler and Cutko, 2010), New Hampshire Natural Heritage Program (Sperduto and Nichols, 2011), and Massachusetts Division of Fisheries and Wildlife (Swain, 2020).
Northern hardwoods dominate this site, particularly white ash, yellow birch, sugar maple and beech. Enriched site indicators (such as blue cohosh and maidenhair fern) are common on this site.
Treethrow and logging are the most common disturbances affecting this site. The site is resilient following these disturbances and succeeds through an herbaceous and shrubby phase prior to tree establishment and eventual return to the reference community. The young forest stands include several species not typically dominant in the reference community, including pin cherry, white birch, aspen, balsam fir, etc.
On gentler slopes, this site may be cultivated for crop or pasture. When cropland or pastureland management ceases, the site either returns to northern hardwoods or may transition to a white pine forest. Once white pine is established, it tends to form a single age stand with low diversity and little understory.
This site includes the following state natural heritage program types:
• Rich Mesic foresst (Sperduto and Nichols 2004)
• Enriched Northern Hardwoods Forest (Gawler and Cutko 2010)
• Sugar Maple Forest (Gawler and Cutko 2010)
• Rich Mesic Forests (Thompson and Sorenson 2000)State and transition model
Custom diagramStandard diagram
More interactive model formats are also available. View Interactive Models
More interactive model formats are also available. View Interactive Models
Click on state and transition labels to scroll to the respective textEcosystem states
State 1 submodel, plant communities
State 2 submodel, plant communities
State 3 submodel, plant communities
State 4 submodel, plant communities
State 1
Reference State / Current PotentialCommunity 1.1
Northern Hardwood ForestMulti-age stand dominated by white ash, yellow birch, sugar maple, and/or beech.
Community 1.2
Herbaceous PhaseWild raspberry, ferns, and other herbs colonize the open land
Community 1.3
Successional ForestDiverse young hardwoods, including species not dominant in the reference community
Community 1.4
Mature Forest 50-80 yr50-80 year old hardwoods. Early successional species (white birch, fir, grey birch, aspen) dying out.
Pathway P1.1-1.2
Community 1.1 to 1.2windthrow, blowdown, fire
Pathway P1.2-1.3
Community 1.2 to 1.3vegetation development (succession)
Key ecosystem services affected
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Genetic resources
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Nutrient cycling
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Primary production
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Climate regulation: carbon cycling and storage
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Soil formation
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Animal biodiversity
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Plant biodiversity
Pathway P1.3-1.4
Community 1.3 to 1.4vegetation development (succession)
Key ecosystem services affected
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Nutrient cycling
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Primary production
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Climate regulation: carbon cycling and storage
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Soil formation
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Animal biodiversity
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Plant biodiversity
Pathway P1.4-1.1
Community 1.4 to 1.1windthrow, blowdown, fire
Key ecosystem services affected
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Genetic resources
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Nutrient cycling
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Climate regulation: carbon cycling and storage
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Wildlife habitat
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Soil formation
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Animal biodiversity
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Plant biodiversity
Pathway P1.4-1.2
Community 1.4 to 1.2windtrhrow, blowdown, fire
Key ecosystem services affected
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Genetic resources
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Nutrient cycling
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Primary production
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Wildlife habitat
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Soil formation
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Animal biodiversity
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Plant biodiversity
State 2
Grassland / Hay landCommunity 2.1
Pasture or Hay LandCleared and planted fields of mostly perennial herbaceous species.
State 3
Crop LandCommunity 3.1
Annual or Perennial CropsCleared and cultivated fields, heavily managed with regular soil disturbance.
State 4
White PineCommunity 4.1
Herbs and ShrubsWild raspberry, ferns, and other herbs colonize the open land
Community 4.2
White Pine ForestSingle age white pine forest.
Pathway P4.1-4.2
Community 4.1 to 4.2Vegetation development (succession)
Key ecosystem services affected
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Aesthetic values
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Genetic resources
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Sense of place
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Nutrient cycling
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Primary production
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Food and fiber: wood products
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Wildlife forage
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Climate regulation: carbon cycling and storage
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Soil formation
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Animal biodiversity
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Plant biodiversity
Pathway P4.2-4.1
Community 4.2 to 4.1harvest, logging
Conservation practices
Forest Stand Improvement Forest Land Management Key drivers
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Timber management
Key ecosystem services affected
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Aesthetic values
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Genetic resources
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Sense of place
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Nutrient cycling
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Primary production
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Food and fiber: wood products
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Wildlife forage
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Climate regulation: carbon cycling and storage
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Soil formation
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Animal biodiversity
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Plant biodiversity
Transition T1-2
State 1 to 2tree removal, pasture or hayfield establishment
Conservation practices
Clearing and Snagging Land Clearing Invasive Plant Species Control Managed Haying/Grazing Key ecosystem services affected
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Aesthetic values
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Genetic resources
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Sense of place
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Nutrient cycling
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Primary production
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Food and fiber: wood products
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Climate regulation: carbon cycling and storage
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Wildlife habitat
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Soil formation
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Animal biodiversity
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Plant biodiversity
Transition T1-3
State 1 to 3Tree clearing, crop establishment
Conservation practices
Clearing and Snagging Cover Crop Land Clearing Key ecosystem services affected
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Aesthetic values
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Genetic resources
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Sense of place
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Nutrient cycling
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Primary production
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Wildlife forage
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Climate regulation: carbon cycling and storage
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Wildlife habitat
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Soil formation
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Animal biodiversity
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Plant biodiversity
Transition T1-4
State 1 to 4selective harvest
Conservation practices
Forest Stand Improvement Forest Land Management Key drivers
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Timber management
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Selective harvest
Key ecosystem services affected
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Aesthetic values
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Genetic resources
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Sense of place
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Nutrient cycling
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Primary production
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Food and fiber: wood products
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Wildlife forage
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Climate regulation: carbon cycling and storage
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Wildlife habitat
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Soil formation
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Animal biodiversity
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Plant biodiversity
Transition R2-1
State 2 to 1abandonment, vegetation development (succession), planting
Conservation practices
Tree/Shrub Site Preparation Tree/Shrub Establishment Upland Wildlife Habitat Management Restoration and Management of Natural Ecosystems Native Plant Community Restoration and Management Invasive Plant Species Control Managed Haying/Grazing Transition T2-4
State 2 to 4tree establishment
Conservation practices
Tree/Shrub Site Preparation Tree/Shrub Establishment Invasive Plant Species Control Key drivers
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Timber management
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Tree planting
Key ecosystem services affected
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Genetic resources
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Sense of place
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Nutrient cycling
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Primary production
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Food and fiber: wood products
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Wildlife forage
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Climate regulation: carbon cycling and storage
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Wildlife habitat
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Soil formation
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Animal biodiversity
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Plant biodiversity
Restoration pathway R3-1
State 3 to 1abandonment, vegetation development (succession), tree planting
Conservation practices
Tree/Shrub Establishment Upland Wildlife Habitat Management Tree/Shrub Pruning Restoration and Management of Natural Ecosystems Native Plant Community Restoration and Management Invasive Plant Species Control Transition T3-4
State 3 to 4tree planting
Conservation practices
Tree/Shrub Site Preparation Tree/Shrub Establishment Invasive Plant Species Control Key drivers
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Timber management
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Tree planting
Key ecosystem services affected
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Aesthetic values
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Genetic resources
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Sense of place
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Nutrient cycling
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Primary production
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Food and fiber: wood products
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Wildlife forage
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Wildlife habitat
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Soil formation
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Animal biodiversity
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Plant biodiversity
Restoration pathway R4-1
State 4 to 1abandonment, vegetation development (succession), plantings
Conservation practices
Tree/Shrub Site Preparation Tree/Shrub Establishment Upland Wildlife Habitat Management Restoration and Management of Natural Ecosystems Native Plant Community Restoration and Management Invasive Plant Species Control Restoration pathway T4-2
State 4 to 2Tree removal, pasture or hay land establishment
Conservation practices
Clearing and Snagging Land Clearing Transition T4-3
State 4 to 3tree removal, cropland establishment
Conservation practices
Clearing and Snagging Cover Crop Land Clearing Additional community tables
Table 5. Community 1.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 6. Community 1.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 7. Community 1.3 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 8. Community 1.4 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 9. Community 2.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 10. Community 3.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 11. Community 4.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 12. Community 4.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Interpretations
Supporting information
Inventory data references
Future work is needed, as described in a future project plan, to validate the information presented in this provisional ecological site description. Future work includes field sampling, data collection and analysis by qualified vegetation ecologists and soil scientists. As warranted, annual reviews of the project plan can be conducted by the Ecological Site Technical Team. A final field review, peer review, quality control, and quality assurance reviews of the ESD are necessary to approve a final document.
Other references
Comer, P., D. Faber-Langendoen, R. Evans, S. Grawler, C. Josse, G. Kittel, S. Menard, M. Pyne, M. Reid, K. Schultz, K. Snow, and J. Teague. 2003. Ecological Systems of the United States: A Working Classification of U.S. Terrestrial Systems. NatureServe, Arlington, Virginia
Edinger, G. J., D. J. Evans, S. Gebauer, T. G. Howard, D. M. Hunt, and A. M. Olivero (editors). 2014. Ecological Communities of New York State. Second Edition. A revised and expanded edition of Carol Reschke’s Ecological Communities of New York State. New York Natural Heritage Program, New York State Department of Environmental Conservation, Albany, NY.
Gawler, S. and A. Cutko. 2010. Natural Landscapes of Maine: A Guide to Natural Communities and Ecosystems. Maine Natural Areas Program, Maine Department of Conservation, Augusta, Maine.
NatureServe. 2021. NatureServe Explorer: An online encyclopedia of life [web application]. NatureServe, Arlington, Virginia. https://explorer.natureserve.org/. (accessed 10 July. 2021).
Soil Survey Staff, Natural Resources Conservation Service, United States Department of Agriculture. 2006. Land Resource Regions and Major Land Resource Areas of the United States, the Caribbean, and the Pacific Basin. Agricultural Handbook 296
Soil Survey Staff, Natural Resources Conservation Service, United States Department of Agriculture. Official Soil Series Descriptions. Available online. (accessed 11 Aug. 2021).
Soil Survey Staff, Natural Resources Conservation Service, United States Department of Agriculture. Soil Climate Research Station Data. Available online. (accessed 23 June. 2021).
Soil Survey Staff, Natural Resources Conservation Service, United States Department of Agriculture. Soil Survey Geographic (SSURGO) Database for [MLRA 141, Maine]. Available online. (accessed 14 Oct. 2021).
Sperduto, D.D. and William F. Nichols. 2011. Natural Communities of New Hampshire. 2nd Ed. NH Natural Heritage Bureau, Concord, NH. Pub. UNH Cooperative Extension, Durham, NH.
Swain, P. C. 2020. Classification of the Natural Communities of Massachusetts. Massachusetts Division of Fisheries and Wildlife, Westborough, MA
USNVC [United States National Vegetation Classification]. 2017. United States National Vegetation Classification Database V2.01. Federal Geographic Data Committee, Vegetation Subcommittee, Washington DC. Available The U.S. National Vegetation Classification (usnvc.org) (accessed 2 July. 2021).Contributors
Christopher Mann
Approval
Nels Barrett, 9/27/2024
Acknowledgments
Nels Barrett and Nick Butler provided considerable review of this ecological site concept.
Rangeland health reference sheet
Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.
Author(s)/participant(s) Contact for lead author Date 06/29/2020 Approved by Approval date Composition (Indicators 10 and 12) based on Annual Production Indicators
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Number and extent of rills:
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Presence of water flow patterns:
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Number and height of erosional pedestals or terracettes:
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Bare ground from Ecological Site Description or other studies (rock, litter, lichen, moss, plant canopy are not bare ground):
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Number of gullies and erosion associated with gullies:
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Extent of wind scoured, blowouts and/or depositional areas:
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Amount of litter movement (describe size and distance expected to travel):
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Soil surface (top few mm) resistance to erosion (stability values are averages - most sites will show a range of values):
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Soil surface structure and SOM content (include type of structure and A-horizon color and thickness):
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Effect of community phase composition (relative proportion of different functional groups) and spatial distribution on infiltration and runoff:
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Presence and thickness of compaction layer (usually none; describe soil profile features which may be mistaken for compaction on this site):
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Functional/Structural Groups (list in order of descending dominance by above-ground annual-production or live foliar cover using symbols: >>, >, = to indicate much greater than, greater than, and equal to):
Dominant:
Sub-dominant:
Other:
Additional:
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Amount of plant mortality and decadence (include which functional groups are expected to show mortality or decadence):
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Average percent litter cover (%) and depth ( in):
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Expected annual annual-production (this is TOTAL above-ground annual-production, not just forage annual-production):
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Potential invasive (including noxious) species (native and non-native). List species which BOTH characterize degraded states and have the potential to become a dominant or co-dominant species on the ecological site if their future establishment and growth is not actively controlled by management interventions. Species that become dominant for only one to several years (e.g., short-term response to drought or wildfire) are not invasive plants. Note that unlike other indicators, we are describing what is NOT expected in the reference state for the ecological site:
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Perennial plant reproductive capability:
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