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Conservation Service
Ecological site PX136X00X720
Basic Upland Forest, Moist
Last updated: 5/02/2025
Accessed: 07/14/2026
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Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.
MLRA notes
Major Land Resource Area (MLRA): 136X–Southern Piedmont
This MLRA is on a large piedmont underlain by metamorphic and igneous bedrock. It stretches from north-central Virginia to east-central Alabama, running parallel to the Appalachian highlands to the northwest and the Atlantic coast to the southeast.
MLRA 136 has only subtle climatic differences with MLRA 148 (Northern Piedmont), with which it shares a common geologic origin. This adjacent MLRA sits to the north. Along the fall line, it shares a boundary with MLRA 133A (Southern Coastal Plain), MLRA 137 (Carolina and Georgia Sand Hills), and 133C (Gulf Coastal Plain). Here, unconsolidated Coastal Plain sediments intersect the much older Piedmont bedrock. Along it's northwestern boundary, it sits adjacent to MLRAs 130B (Southern Blue Ridge), 130A (Northern Blue Ridge), and 128 (Southern Appalachian Ridges and Valleys). These MLRAs are distinguished from the Southern Piedmont by topographic and elevational differences, as well as differences in the age, origin, and degree of metamorphism of the underlying bedrock.
Five states are intersected by the MLRA, including North Carolina (29 percent), Georgia (27 percent), Virginia (20 percent), South Carolina (17 percent), and Alabama (7 percent). The MLRA extent makes up about 63,720 square miles (165,034 square kilometers).
MLRA PHYSIOGRAPHY
The landscape is generally rolling to hilly, with a well-defined drainage pattern. Streams have dissected the original Piedmont plateau, forming narrow ridgetops, somewhat broad interfluves, and short, steep side slopes adjacent to the streams and drainageways. With some exceptions, the valley floors are generally narrow and make up about 10 percent or less of the land area. The associated stream terraces are generally small and of minor extent.
The landscape is moderately dissected overall, with isolated erosional remnants (monadnocks) and other areas of high topographic relief interspersed. Over most of the MLRA, elevation ranges from approximately 325 to 1,315 feet (100 to 400 meters), with elevations generally increasing toward the Appalachian Highlands, in the upper Piedmont, and decreasing toward the Coastal Plain, in the lower Piedmont.
The major rivers that cross this area en route to the ocean include, from north to south, the James, Roanoke, Cape Fear, Savannah, Altamaha, Chattahoochee, and Alabama Rivers. These rivers typically originate within the Piedmont or in the Blue Ridge. They flow east and south across the Coastal Plain and empty into the Atlantic Ocean or the Gulf of America.
MLRA GEOLOGY
Precambrian and Paleozoic metamorphic and igneous rocks underlie almost all of this MLRA. The dominant metamorphic rock types include gneiss, schist, slate, argillite, and phyllite, among others. Dominant igneous rock types include granite and other related felsic crystalline rocks. Mafic intrusive rocks, including gabbro, diabase, amphibolite, and other dark colored rocks, underlie a minority of the upland landscape. These mafic intrusions crop out in the form of dikes and sills, and often weather to produce soils high in base cations.
The Carolina Slate Belt runs lengthwise through the east-central part of the MLRA, in southern Virginia, North Carolina, South Carolina, and the eastern-most part of the Georgia Piedmont. This region is underlain by fine-grained metasedimentary and metavolcanic rock, which generally weathers to produce soils high in silt.
From Virginia to North Carolina, and in a single county in South Carolina, fault-bounded Triassic Basins are scattered amongst the igneous and metamorphic uplands. These basins are underlain by Triassic and Jurassic siltstone, shale, sandstone, and mudstone, which were laid down in response to continental rifting and subsequent erosion during the Mesozoic era.
MLRA SOILS
The dominant soil orders of the MLRA are Ultisols, Inceptisols, and Alfisols. Ultisols and Alfisols are typically found on more stable landforms, such as interfluves, gentle hillslopes, broad ridgetops, and stream terraces, while Inceptisols are typically found on less stable landforms, including flood plains, steep hillslopes, and narrow ridgetops.
Soils of the region predominantly have a thermic temperature regime, a udic moisture regime, and generally have kaolinitic or mixed mineralogy. In the upper Piedmont of Virginia and North Carolina however, soils have a mesic soil temperature regime, as depicted in figure 2. The mesic soil temperature regime portion of the MLRA is oriented from northeast to southwest and occupies approximately 18 percent of the MLRA extent, or 11,729 square miles (30,377 square kilometers).
Broadly speaking, soils of the Southern Piedmont uplands are shallow to very deep, well drained, and loamy or clayey. Soils of the river valleys are generally very deep, well to poorly drained, and loamy. Soils tend to be finer-textured than in Coastal Plain regions.
MLRA CLIMATE
In general, precipitation is evenly distributed throughout the year in this MLRA, with occasional drought-like conditions extending from late summer into autumn. During the growing season, most of the rainfall comes from high-intensity, convective thunderstorms. Significant moisture also comes from the movement of warm and cold fronts across the MLRA from November to April. High amounts of rain can also occur during hurricanes, usually during the months of August through October.
Over most of the MLRA, snowfall is typically light, though overall, the mesic soil temperature regime portion of the MLRA features colder temperatures, more snowfall, and a shorter growing season than in the thermic portion. The cooler climate in this region supports an increase in species with northern or Blue Ridge affinities. Both the mean annual temperature and the length of the freeze-free period increase from north to south and with decreasing elevation from the upper to the lower Piedmont.
MLRA LAND USE AND RESOURCES
Once largely cultivated, much of this region is now planted to loblolly pine or has reverted to successional pine and hardwood forests. The more productive lands support small to medium-size family farms that produce crops and livestock, while the less productive lands have been in forest for some time. Most of the open areas are used for grazing beef cattle, though in years past, dairy cattle were also important to the local economy. The principal crops of the region include corn, soybeans, and small grains. Burley tobacco remains a crop of local importance. Cotton is grown in the thermic soil temperature regime portion of the MLRA.
Several major land cover transformations have occurred in the Southern Piedmont over the past several centuries; from open woodlands sculpted by fire, to farmland, to closed forests and planted pine, past land uses have played an outsized role in shaping present-day soils and vegetation patterns in the region. Land-use intensity peaked with the arrival of the industrial revolution, which gradually increased demand for textiles. Cotton became the dominant crop over much of the region.
In spite of early successes, two centuries of poor management practices accelerated soil erosion, stripping away the fertility and moisture-supplying capacity of soils. In addition to soil losses in the uplands, legacy sediments derived from the eroded land rapidly accumulated in the river valleys below, often leading to changes in hydrology and flooding frequency.
After being stripped of it's loamy topsoil, many areas of the Piedmont had been so badly eroded as to render the land unsuitable or economically impractical for agriculture. The effects of erosion were widespread, with cumulative soil loss estimates ranging from 5 to 10 inches on average. The steeper slopes, which had often been cleared and farmed at the height of the Cotton era, generally suffered greater losses. By the 1930's, crop production was in rapid decline in the Southern Piedmont. The loss of soil productivity due to erosion, losses to the cotton boll weevil, development of synthetic fibers, and the onset of the Great Depression all contributed to rapid abandonment of cropland. By 1960, cropland acres had decreased by more than 50 percent in nearly every county in the Southern Piedmont.
While crop production is still important today on the more productive lands, those of lower productivity, or those that were subject to severe erosion, were often abandoned some time ago. Typically, they have either reverted to forest, or have been converted to other uses. Although the productivity of soils was greatly reduced through erosion, less intensive land uses such as grazing and forestry were still feasible. These land uses gained popularity as patterns of urban migration, low commodity prices, and other factors gradually made crop production less economical on the marginal lands.
In recent years, large-scale adoption of soil conservation practices have led to better outcomes with respect to erosion in much of MLRA, increasing the economic viability and long-term sustainability of Piedmont farms. Despite some success, water erosion remains one of the most important soil resource concerns in the MLRA.
Other major resource concerns include increasing conversion of prime farmland and farmland of statewide importance to urban uses. Throughout the MLRA, metropolitan areas are expanding into lands that have historically been used for timber or agriculture. This change in land use is occurring rapidly in the corridor called the Piedmont Crescent, which extends from Atlanta, Georgia, to Raleigh, North Carolina.
HISTORIC VEGETATION COVER
Over most of the Southern Piedmont uplands, the historic oak-hickory, or oak-hickory-pine forest, once covered large portions of the landscape. It was dominated by upland oaks, such as white oak (Quercus alba), northern red oak (Quercus rubra), and southern red oak (Quercus falcata), with a smaller contribution from hickories (Carya spp.) and pines. The principal pine species are shortleaf pine (Pinus echinata), loblolly pine (Pinus taeda), and to the north and west, Virginia pine (Pinus virginiana). In the southernmost and easternmost portions of the MLRA, the historic montane longleaf pine forest, dominated by longleaf pine (Pinus palustris), shortleaf pine (P. echinata), and dry-site oaks, was found on ridgetops and steep south or west-facing slopes.
According to historic accounts, forests and woodlands of the past were generally more open and park-like, having been exposed to a more frequent fire regime. Piedmont prairies, likely maintained by Native Americans, were also reportedly common across the landscape, as were fire-maintained canebrakes along the streams (Trimble 1974; Daniels 1987; Griffith et al. 2002; Van Lear et al. 2004; Dearman and James 2019; Schomberg et al. 2020; USDA-NRCS 2022).LRU notes
MLRA 136 is one of the largest MLRAs in the United States. It has a broad north-south and east-west extent and covers a wide range of elevations. The MLRA is partitioned by the mesic-thermic line, which divides the MLRA into mesic and thermic soil temperature regimes (figure 2.). The mesic soil temperature regime was delineated based on estimates of the native range of loblolly pine, which was historically absent in this part of the MLRA. In addition, this region is said to represent the northern and western limits of cotton production, an important crop to the south and east.
ESDs developed for this MLRA were split geographically into mesic and thermic ecological site concepts. Climate variation across the MLRA extent warrants the development of Land Resource Unit (LRU) classifications, to further subdivide the MLRA and support more precise Ecological Site Descriptions.Classification relationships
APPLICABLE USNVC ASSOCIATIONS
CEGL007232 Quercus alba - Quercus rubra - Carya (ovata, carolinae-septentrionalis) / Cercis canadensis
APPLICABLE EPA ECOREGIONS
Level III: 45. Piedmont
Level IV: 45a. Southern Inner Piedmont; 45b. Southern Outer Piedmont; 45c. Carolina Slate Belt; 45f. Northern Outer Piedmont; 45g. Triassic Basins; 45i. Kings Mountain; 45d. Talladega Upland; 45h. Pine Mountain Ridges (EPA 2013).
APPLICABLE USFS ECOLOGICAL UNITS
Domain: Humid Temperate
Division: Subtropical
Ecological province: 231. Southeastern Mixed Forest
Ecological sections: 231I.Central Appalachian Piedmont; 231A. Southern Appalachian Piedmont (Cleland et al. 2007).
Based on the USGS physiographic classification system (Fenneman and Johnson 1946), most of MLRA 136 is in the Piedmont Upland section of the Piedmont province, in the Appalachian Highlands division.Ecological site concept
This ecological site includes moist uplands that are rich in base cations, on broad ridges, interfluves, and hillslopes. It is geographically restricted to the thermic soil temperature regime portion of the MLRA.
Soils on this ecological site are weathered from mafic intrusive rock. They are typically deep to very deep, well drained Alfisols, which formed in residuum from dark colored rocks high in ferromagnesian minerals. Base saturation is greater than or equal to 35 percent in the subsoil.
The reference state supports a closed to somewhat open canopy oak-hickory forest dominated by white oak (Quercus alba), northern red oak (Quercus rubra), and other upland oaks, with a higher proportion of hickories, elms, and ashes, and greater species diversity than is typical of similar acidic forest types. Dominant land uses include cropland, pasture and hayland, planted pine, and various urban or suburban uses.
ES CHARACTERISTICS SUMMARY
• Thermic soil temperature regime
• Occurs on Piedmont uplands, on broad ridges, interfluves, and hillslopes
• Parent materials: mafic intrusive rock
• Base saturation: ≥ 35 percent in the subsoil
• Seasonal high water table: usually absent within 72 inches (though the minimum allowable depth is ≥ 40 inches from the soil surface)
• Depth to bedrock: ≥ 40 inches, usually deeper
• The available water storage capacity of the profile (from the soil surface to 80 inches, or to paralithic or lithic bedrock, whichever is shallower) is greater than or equal to 6 inches
• Soils: deep to very deep, well drained AlfisolsAssociated sites
PX136X00X710 Basic Upland Woodland, Expansive Clay, Seasonally Wet and Dry
Found in similar or slightly lower landscape positions. The seasonal high water table is shallower (12-40 inches from the soil surface). The clay fraction is dominated by shrink-swell clay minerals, producing extremes in plant available water over the growing season, reducing cover from moisture-loving plant species such as northern red oak (Quercus rubra).
PX136X00X730 Basic Upland Forest, Depth Restriction, Dry
Generally found in similar or slightly higher landscape positions, on less stable and more steeply sloping parts of the landscape where soils are shallower to bedrock (< 40 inches). Soils are more droughty and support a relative increase in dry-site species.
PX136X00X820 Acidic Upland Forest, Moist
Widespread on the surrounding acidic uplands. It is underlain either by acidic igneous or metamorphic rock, or by highly weathered mafic materials that have been leached of base cations. Base saturation is less than 35 percent. This ecological site supports similar species in the canopy, as the relative moisture status is similar, but basic indicator species are generally scarce or absent in the understory.
Similar sites
PX136X00X220 Mesic Temperature Regime, Basic Upland Forest, Moist
The soil temperature regime is mesic, occurring outside of the native range of loblolly pine (Pinus taeda).
PX136X00X820 Acidic Upland Forest, Moist
The relative soil moisture status is similar, but soils are more thoroughly leached of base cations. Base saturation is less than 35 percent in the subsoil. This ecological site supports similar species in the canopy, but basic indicator species are generally scarce or absent in the understory.
Figure 1. EPA level IV ecoregions of the Southern Piedmont (45).
Figure 2. Spatial illustration of soil temperature regimes of the Southern Piedmont.
Figure 3. Spatial extent of this ecological site representing the major areas where this site is important on the landscape.
Table 4. Dominant plant species
Tree (1) Quercus alba
(2) CaryaShrub (1) Cercis canadensis
(2) ViburnumHerbaceous (1) Desmodium nudiflorum
(2) Maianthemum racemosum ssp. racemosumLegacy ID
F136XY720NC
Physiographic features
This ecological site is found on broad ridges, interfluves, and hillslopes. It is found throughout the thermic soil temperature regime portion of the Piedmont, in virtually every ecoregion in this part of the MLRA. Representative locations are gently sloping to moderately steep, with a representative slope of 2 to 10 percent and a maximum slope of 25 percent.
The geologic substrate on this ecological site is mafic intrusive rock, such as diabase, gabbro, and other related dark colored rocks high in ferromagnesian minerals. The material was transported upward through fissures in the earth's crust in the form of mafic dikes and sills. These structures cut through the surrounding country rock. As a result, this ecological site often occurs in small patches, and is often adjacent to or intermingled with acidic uplands, or basic uplands with dissimilar soil moisture conditions.
Figure 1. Typical soil-landscape relationships of the Southern Piedmont Mixed Felsic and Mafic Soil System. Enon soils are associated with this ecological site, depicted here on interfluves.
Figure 2. Typical soil-landscape relationships of the Southern Piedmont Mixed Felsic and Mafic Soil System. Mecklenburg soils are associated with this ecological site, depicted here on interfluves.
Table 5. Representative physiographic features
Hillslope profile (1) Summit
(2) Backslope
(3) Shoulder
Landforms (1) Piedmont > Interfluve
(2) Piedmont > Ridge
(3) Piedmont > Hillslope
Runoff class Low to high Flooding frequency None Ponding frequency None Elevation 370 – 980 ft Slope 2 – 10 % Water table depth 72 – 999 in Aspect Aspect is not a significant factor Table 6. Representative physiographic features (actual ranges)
Runoff class Negligible to very high Flooding frequency None Ponding frequency None Elevation 210 – 1090 ft Slope 2 – 25 % Water table depth 72 – 999 in Climatic features
On this ecological site, the average mean annual precipitation is 48 inches. On average, the rainiest months occur from July through August, as well as in March. The driest months occur from April through May, along with October.
Table 7 Representative climatic features
Frost-free period (characteristic range) 160-190 days Freeze-free period (characteristic range) 190-230 days Precipitation total (characteristic range) 50-50 in Frost-free period (actual range) 160-200 days Freeze-free period (actual range) 180-250 days Precipitation total (actual range) 40-60 in Frost-free period (average) 180 days Freeze-free period (average) 210 days Precipitation total (average) 50 in Characteristic rangeActual rangeBarLineFigure 3. Monthly precipitation range
Characteristic rangeActual rangeBarLineFigure 4. Monthly minimum temperature range
Characteristic rangeActual rangeBarLineFigure 5. Monthly maximum temperature range
BarLineFigure 6. Monthly average minimum and maximum temperature
Figure 7. Annual precipitation pattern
Figure 8 Annual average temperature pattern
Climate stations used
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(1) GREENBAY 3 NE [USC00443565], Burkeville, VA
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(2) CHARLOTTE COURT HOUSE [USC00441585], Keysville, VA
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(3) CAMP PICKETT [USC00441322], Blackstone, VA
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(4) LAWRENCEVILLE 3 E [USC00444768], Freeman, VA
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(5) CLARKSVILLE [USC00441746], Clarksville, VA
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(6) JOHN H KERR DAM [USC00444414], Boydton, VA
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(7) ARCOLA [USC00310241], Macon, NC
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(8) ROXBORO 7 ESE [USC00317516], Roxboro, NC
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(9) SANFORD 8 NE [USC00317656], Sanford, NC
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(10) CHAPEL HILL WILLIAMS AP [USW00093785], Chapel Hill, NC
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(11) OXFORD AG [USC00316510], Oxford, NC
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(12) FOREST CITY 6 SW [USC00313150], Forest City, NC
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(13) GASTONIA MUNI AP [USW00053870], Gastonia, NC
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(14) MONROE 2 SE [USC00315771], Monroe, NC
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(15) WADESBORO [USC00318964], Wadesboro, NC
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(16) ROCK HILL YORK CO AP [USW00053871], Rock Hill, SC
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(17) KERSHAW 1SW [USC00384690], Kershaw, SC
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(18) CEDAR CREEK 2E [USC00381479], Blythewood, SC
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(19) SALUDA [USC00387631], Saluda, SC
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(20) CLEMSON OCONEE CO AP [USW00053850], Seneca, SC
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(21) WALHALLA [USC00388887], Walhalla, SC
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(22) GREENVILLE DWTN AP [USW00013886], Greenville, SC
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(23) NINETY NINE ISLANDS [USC00386293], Blacksburg, SC
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(24) CALHOUN FALLS [USC00381277], Calhoun Falls, SC
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(25) MC CORMICK 9 E [USC00385658], Mc Cormick, SC
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(26) PICKENS [USC00386831], Pickens, SC
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(27) WASHINGTON 2 ESE [USC00099157], Washington, GA
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(28) CUMMING 2N [USC00092408], Cumming, GA
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(29) MULBERRY GROVE [USC00096148], Smiths Station, GA
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(30) DAHLONEGA [USC00092475], Dahlonega, GA
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(31) JONESBORO [USC00094700], Jonesboro, GA
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(32) LA GRANGE 1N [USC00094949], LaGrange, GA
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(33) OPELIKA [USC00016129], Opelika, AL
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(34) LAFAYETTE 2W [USC00014502], Lafayette, AL
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(35) ROCK MILLS [USC00017025], Roanoke, AL
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(36) ALEXANDER CITY [USC00010160], Alexander City, AL
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(37) EXPERIMENT [USC00093271], Griffin, GA
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(38) ASHEBORO 2 W [USC00310286], Asheboro, NC
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(39) CHESNEE 7 WSW [USC00381625], Chesnee, SC
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(40) CLEMSON UNIV [USC00381770], Clemson, SC
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(41) GREENWOOD [USC00383754], Greenwood, SC
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(42) ATHENS BEN EPPS AP [USW00013873], Athens, GA
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(43) ASHLAND 3 ENE [USC00010369], Ashland, AL
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(44) ROCKFORD 3 ESE [USC00017020], Rockford, AL
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(45) GAINESVILLE [USC00093621], Gainesville, GA
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(46) MILLEDGEVILLE [USC00095874], Milledgeville, GA
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(47) WEST POINT [USC00099291], Lanett, GA
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(48) SIMMS WTP [USC00387885], Chesnee, SC
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(49) CHARLOTTE DOUGLAS AP [USW00013881], Charlotte, NC
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(50) CARROLLTON [USC00091640], Carrollton, GA
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(51) COVINGTON [USC00092318], Covington, GA
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(52) ALBEMARLE [USC00310090], Albemarle, NC
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(53) NEWBERRY [USC00386209], Newberry, SC
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(54) COLUMBUS METRO AP [USW00093842], Columbus, GA
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(55) DALLAS 7 NE [USC00092485], Dallas, GA
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(56) SILER CITY 2 N [USC00317924], Siler City, NC
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(57) CHASE CITY [USC00441606], Chase City, VA
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(58) CROZIER [USC00442142], Maidens, VA
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(59) AMELIA 8 NE [USC00440188], Amelia Court House, VA
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(60) AMELIA COURTHOUSE 1 [USC00440187], Amelia Court House, VA
">Influencing water features
This ecological site is not influenced by surface or ground water features.
Soil features
Soils on this ecological site are typically deep to very deep, well drained Alfisols. The available water storage capacity of the profile is generally greater than 6 inches. Unweathered or partially weathered bedrock may be found at depths greater than 40 inches of the surface, but most soils associated with this ecological site are deeper to bedrock. Typically, parent materials are residuum derived from mafic intrusive rock, such as diabase, gabbro, amphibolite, and other related dark colored rocks high in ferromagnesian minerals.
Reaction in the subsoil is typically strongly acid to neutral (pH 5.1 to 7.3). In the surface layers, reaction varies with land use and management. Under low input or forested conditions, it generally falls somewhere between pH 5.1 and 6.5. Base saturation is greater than or equal to 35 percent in the subsoil. These soils are typically in a fine particle size family.
Soils on this ecological site have a thermic soil temperature regime, which is characterized by a mean annual soil temperature of 15°C to 22°C and a winter to summer temperature differential of 6°C or more in the subsoil.
Modal taxa include: Typic Hapludalfs, Ultic Hapludalfs
Modal soil series include: Winnsboro, Enon, Mecklenburg
Other soils attributed to this ecological site include Coronaca, Gundy, Rowan, and Skyuka.
Figure 9. An illustration of a soil profile belonging to the Enon series, a representative soil series associated with this ecological site.
Figure 10. A soil profile of the Winnsboro series, a representative soil series associated with this ecological site.
Figure 11. A soil profile of the Mecklenburg series, a representative soil series associated with this ecological site.
Table 8. Representative soil features
Parent material (1) Residuum – diabase
(2) Residuum – gabbro
(3) Residuum – amphibolite
(4) Residuum – diorite
Surface texture (1) Sandy loam
(2) Fine sandy loam
(3) Loam
(4) Clay loam
(5) Sandy clay loam
Family particle size (1) Fine
Drainage class Well drained Permeability class Moderately slow to moderate Depth to restrictive layer 67 – 999 in Soil depth 67 – 80 in Surface fragment cover <=3" Not specified Surface fragment cover >3" Not specified Available water capacity
(0-80in)8 – 11 in Soil reaction (1:1 water)
(0-10in)5.1 – 6.5 Subsurface fragment volume <=3"
(0-80in)0 – 10 % Subsurface fragment volume >3"
(0-80in)0 – 2 % Table 9. Representative soil features (actual values)
Drainage class Well drained Permeability class Moderately slow to moderately rapid Depth to restrictive layer 59 – 999 in Soil depth 59 – 80 in Surface fragment cover <=3" 0 % Surface fragment cover >3" 0 % Available water capacity
(0-80in)6 – 13 in Soil reaction (1:1 water)
(0-10in)5.1 – 7.3 Subsurface fragment volume <=3"
(0-80in)0 – 16 % Subsurface fragment volume >3"
(0-80in)0 – 3 % Ecological dynamics
U.S. National Vegetation Classification (USNVC) associations that are consistent with reference conditions on this ecological site include CEGL007232 Quercus alba - Quercus rubra - Carya (ovata, carolinae-septentrionalis) / Cercis canadensis. This concept is closely related to the reference community (USNVC 2022).
MATURE FORESTS
The reference state supports a closed to somewhat open canopy oak-hickory forest, with a higher proportion of hickories, elms, and ashes, and greater species diversity than is typical of similar acidic forest types. Acid-loving species are usually scarce or absent in the understory and basic indicator species are typically present in greater numbers.
In the reference state, the canopy is dominated by mesophytic and dry-mesophytic oaks. Dominant canopy species include white oak (Quercus alba), northern red oak (Quercus rubra), and various hickories (Carya ovata, C. carolinae-septentrionalis, C. tomentosa, C. glabra, C. ovalis, etc.). Unlike drier basic upland forests, dry-site oaks (such as Quercus stellata, Q. marilandica, and Q. velutina) are usually of low cover in the canopy.
Other canopy species which are characteristic, but generally of lower cover include winged elm (Ulmus alata), American elm (Ulmus americana), and black walnut (Juglans nigra). Tuliptree (Liriodendron tulipifera), though more abundant in the early stages of succession, is also characteristic of mature stands. It is usually of very low cover however, colonizing and reproducing chiefly in canopy gaps.
In the reference state, the subcanopy layer includes basic indicator species such as eastern redbud (Cercis canadensis), hophornbeam (Ostrya virginiana), white ash (Fraxinus americana), and southern sugar maple (Acer floridanum). Subcanopy generalists include flowering dogwood (Cornus florida), red maple (Acer rubrum), American beech (Fagus grandifolia), black cherry (Prunus serotina), and white fringetree (Chionanthus virginicus).
In the shrub layer, characteristic basic indicator species include downy arrowood (Viburnum rafinesqueanum), blackhaw (Viburnum prunifolium), and coralberry (Symphoricarpos orbiculatus). On moist lower slopes, species more commonly associated with flood plains often make an appearance, including northern spicebush (Lindera benzoin), painted buckeye (Aesculus sylvatica), and American bladdernut (Staphylea trifolia), along with other mesophytic species such as eastern sweetshrub (Calycanthus floridus). Generalists frequent in the shrub layer include bursting-heart (Euonymus americanus) and several ubiquitous vines.
Under reference conditions, the herb layer can be highly diverse, including a suite of moist-site basic indicator species and several species of conservation concern. On the best examples, especially where fire has been reintroduced, the herb layer is well-developed, with up to 50 percent or more total cover and impressive species richness. In many locations however, large deer populations impede the development the herb layer, producing lower than expected species richness. Species richness can be increased through effective deer population management.
Basic indicator species typical of the herb layer include bearded shorthusk (Brachyelytrum erectum), twoflower melicgrass (Melica mutica), hairy woodland brome (Bromus pubescens), and several skullcap species (Scutellaria spp.). Generalists typical of the herb layer include nakedflower ticktrefoil (Desmodium nudiflorum), feathery false lily of the valley (Maianthemum racemosum ssp. racemosum), ebony spleenwort (Asplenium platyneuron), licorice bedstraw (Galium circaezans), and Virginia snakeroot (Aristolochia serpentaria).
On moist lower slopes, species requiring circumneutral soils and sheltered, continually moist conditions make an appearance, including Canadian wildginger (Asarum canadense), richweed (Collinsonia canadensis), black baneberry (Actaea racemosa), northern maidenhair (Adiantum pedatum), heartleaf foamflower (Tiarella cordifolia), and many others.
DYNAMICS OF NATURAL SUCCESSION AND FIRE ECOLOGY
On Piedmont uplands, the historical influence of fire on successional dynamics was likely expressed on a continuum, from dry to moist, where moist or sheltered sites were shaped more by gap-driven dynamics and dry or exposed sites more by fire. On intermediate sites, their respective influence on successional dynamics probably fell somewhere in between. While the historic fire return interval is thought to be relatively similar across most of the Southern Piedmont uplands, moister sites were less prone to fire and hence burned less completely and at lower intensities than drier sites.
Like other moist oak-hickory forests in the region, successional dynamics are thought to be primarily gap-driven, with small-scale natural disturbances such as windthrow, drought, and disease, usually affecting only small portions of the forest at a time. Canopy gaps are readily colonized by early successional herbs and shrubs, and later by pines and opportunistic hardwoods. These localized events are inconspicuous, but cumulatively they help shape the age class distribution, structure, and species composition in these forests.
In the past, regular low-intensity fires would have kept the understory somewhat more open than at present and constrained the growth of fire-intolerant woody species. Periodic severe fires would have likely occurred during unusually dry and windy conditions, presumably resulting in catastrophic tree mortality and stand replacing changes. The reduction in the frequency of fires over the past century has allowed shade-tolerant, fire-sensitive trees such as red maple (Acer rubrum), American beech (Fagus grandifolia), and American holly (Ilex opaca) to become more abundant in many upland forests in the Southeast.
A combination of prescribed burns and selective removals can open up the understory and constrain the growth of fire-intolerant opportunistic species, thereby restoring the health and vigor of forests that evolved under a more regular fire regime.
YOUNG SECONDARY FORESTS
On relatively undisturbed sites, stands are uneven-aged, with at least some old trees present. In areas that were cultivated in the recent past, even-aged pine stands dominate the landscape, being replaced by oaks and hickories only as the pines die.
In general, young secondary forests on this ecological site are dominated by loblolly and shortleaf pines, along with opportunistic hardwoods such as sweetgum (Liquidambar styraciflua), red maple (Acer rubrum), and tuliptree (Liriodendron tulipifera). Oaks and hickories, including white oak (Q. alba), northern red oak (Q. rubra), and southern red oak (Q. falcata), along with hickories (Carya spp.), are usually confined to the understory of young secondary stands. Their growth is temporarily suppressed by the cover of faster growing tree species.
In the central Piedmont of North Carolina and Virginia, Virginia pine (P. virginiana) becomes increasingly important in young secondary stands, particularly in the Carolina Slate Belt and even more so in regions to the west or north. In regions further south, loblolly pine is typically the more competitive pioneer under most site conditions, apart from higher elevation areas of the upper Piedmont where Virginia pine becomes more abundant (Wharton 1978; Barry 1980; Peet and Christensen 1980, 1987; Skeen et al. 1980; Nelson 1986; Schafale and Weakley 1990; Oakley et al. 1995; Cowell 1998; Spira 2011; Fleming 2012; Guyette et al. 2012; Schafale 2012a, 2012b; Edwards et al. 2013; Fleming et al. 2021; Spooner et al. 2021; Greenberg et al. 2021).
SPECIES LIST
Canopy layer: Quercus alba, Quercus rubra, Carya ovata, Carya carolinae-septentrionalis, Carya glabra, Carya tomentosa, Carya ovalis, Fraxinus americana, Ulmus alata, Ulmus americana, Juglans nigra, Quercus shumardii, Liriodendron tulipifera, Pinus echinata, Pinus taeda, Quercus falcata, Carya illinoinensis,
Subcanopy layer: Cercis canadensis, Ostrya virginiana, Cornus florida, Acer floridanum, Fraxinus americana, Acer rubrum, Fagus grandifolia, Ilex opaca, Prunus serotina, Carya spp., Ulmus rubra, Diospyros virginiana, Juniperus virginiana, Chionanthus virginicus, Frangula caroliniana, Celtis spp., Tilia americana, Magnolia tripetala,
Shrub layer: Viburnum rafinesqueanum, Viburnum prunifolium, Symphoricarpos orbiculatus, Viburnum rufidulum, Euonymus americanus, Calycanthus floridus, Viburnum acerifolium, Aesculus sylvatica, Aesculus pavia, Aesculus parviflora, Lindera benzoin, Corylus americana, Staphylea trifolia, Philadelphus inodorus
Vines/lianas: Vitis rotundifolia, Parthenocissus quinquefolia, Toxicodendron radicans, Lonicera sempervirens, Bignonia capreolata, Matelea carolinensis, Smilax herbacea, Loncera japonica (I),
Herb layer - forbs: Desmodium nudiflorum, Maianthemum racemosum ssp. racemosum, Asplenium platyneuron, Erythronium umbilicatum, Galium circaezans, Polygonatum biflorum, Solidago caesia, Aristolochia serpentaria, Scutellaria elliptica, Scutellaria integrifolia, Geranium maculatum, Polystichum acrostichoides, Asarum canadense, Galium triflorum, Smallanthus uvedalius, Sanguinaria canadensis, Collinsonia canadensis, Podophyllum peltatum, Cynoglossum virginianum, Uvularia perfoliata, Hypoxis hirsuta, Ranunculus hispidus, Actaea racemosa, Tiarella cordifolia, Heuchera americana, Adiantum pedatum, Phryma leptostachya, Sisyrinchium mucronatum, Cardamine concatenata, Ligusticum canadense, Hepatica nobilis var. obtusa, Agrimonia pubescens, Agrimonia rostellata, Scutellaria ovata, Scutellaria serrata, Viola triloba var. triloba, Hybanthus concolor, Phegopteris hexagonoptera, Botrychium virginianum, Erythronium americanum, Desmodium cuspidatum,
Herb layer - graminoids: Dichanthelium spp., Carex spp. (laxiflora, rosea, blanda, striatula, retroflexa, planispicata, kraliana), Melica mutica, Dichanthelium boscii, Festuca subverticillata, Bromus pubescens, Elymus hystrix, Chasmanthium sessiliflorum, Brachyelytrum erectum,
(I) = introducedState and transition model
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Click on state and transition labels to scroll to the respective textEcosystem states
States 1, 5 and 6 (additional transitions)
States 2, 5 and 6 (additional transitions)
T1A - Clearcut logging or other large-scale disturbances that cause canopy removal. T1B - Selective removals of the most valuable timber specimens, leaving inferior trees behind. T1C - Mechanical tree/brush/stump/debris removal, seedbed preparation, and planting of perennial grasses and forbs. T1D - Mechanical tree/brush/stump/debris removal, seedbed preparation, applications of fertilizer/lime, and planting of crop or cover crop seed. T2A - Long-term natural succession. T2B - Site preparation and tree planting. T2C - Mechanical tree/brush/stump/debris removal, seedbed preparation, and planting of perennial grasses and forbs. T2D - Mechanical tree/brush/stump/debris removal, seedbed preparation, applications of fertilizer/lime, weed control, planting of crop or cover crop seed. T3A - Clearcut logging or other large-scale disturbances that cause canopy removal. T3C - Mechanical tree/brush/stump/debris removal, seedbed preparation, applications of fertilizer/lime, weed control, and planting of perennial grasses and forbs. T3D - Mechanical tree/brush/stump/debris removal, seedbed preparation, applications of fertilizer/lime, weed control, planting of crop or cover crop seed. T4A - Abandonment of forestry practices. T4B - Timber harvest, mechanical stump and debris removal, seedbed preparation, and planting of perennial grasses and forbs. T4C - Timber harvest, mechanical stump and debris removal, seedbed preparation, fertilizer/lime, weed control, planting of crop or cover crop seed. T5A - Long-term cessation of grazing. T5B - Site preparation and tree planting. T5C - Seedbed preparation, applications of fertilizer/lime, weed control, and planting of crop or cover crop seed. T6A - Agricultural abandonment. T6B - Site preparation and tree planting. T6C - Seedbed preparation, weed control, and planting of perennial grasses and forbs. State 1 submodel, plant communities
1.1A - Long-term exclusion of fire. 1.2A - Prescribed burns and selective removals. State 2 submodel, plant communities
2.1A - Clearcut logging. 2.2A - Natural succession. 2.2B - Brush management. 2.3A - Natural succession. State 6 submodel, plant communities
6.1A - Conventional tillage is reintroduced. 6.2A - Implementation of conservation tillage and other soil conservation practices State 1
Reference StateThis mature forest state is generally dominated by mesophytic and dry-mesophytic oaks. It usually has a greater proportion of hickories, elms, and ashes than acidic oak-hickory forest types.
Characteristics and indicators. Stands are uneven-aged with at least some old trees present. Pines make up a minority of the canopy cover.
Resilience management. Deer population management is critical to sustaining the diversity of herbaceous understory species.
Community 1.1
Moist Basic Oak-Hickory Forest - Fire Maintained PhaseThis is a closed to somewhat open canopy mature forest community/phase. Regular low-intensity fires have been reintroduced, keeping the understory somewhat open, increasing the cover and diversity of herbaceous species and limiting the importance of fire-intolerant woody species.
Resilience management. This community/phase is maintained through regular prescribed burns. The recruitment of fire-adapted oaks and pines benefits from regular low-intensity ground fires, as these forests evolved under a more regular fire regime. Tree ring data suggests that the mean fire return interval of the past in the Southern Piedmont is approximately 6 years, though the actual return interval varied from 3 to 16 years. To approximate the pre-colonial fire regime, prescribed burns should be carried out every 4 to 8 years.
Forest overstory.The overstory is dominated by oaks. Representative species include white oak (Quercus alba) and northern red oak (Quercus rubra), with a smaller but significant contribution from hickories. Pines are typically scattered throughout the forest. Characteristic hickory species include shagbark hickory (Carya ovata), southern shagbark hickory (C. carolinae-septentrionalis), and red hickory (C. ovalis), along with the more widespread pignut and mockernut hickories (C. glabra, C. tomentosa). Shortleaf pine (Pinus echinata) is representative of the pine component.
Forest understory. Characteristic understory tree species include eastern redbud (Cercis canadensis), flowering dogwood (Cornus florida), winged elm (Ulmus alata), white ash (Fraxinus americana), white fringetree (Chionanthus virginicus), and several species of hickory (Carya spp.).<br /> <br /> Characteristic understory shrub species include coralberry (Symphoricarpos orbiculatus), blackhaw (Viburnum prunifolium), and eastern sweetshrub (Calycanthus floridus).
Dominant plant species
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white oak (Quercus alba), tree
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northern red oak (Quercus rubra), tree
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shagbark hickory (Carya ovata), tree
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southern red oak (Quercus falcata), tree
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eastern redbud (Cercis canadensis), tree
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flowering dogwood (Cornus florida), tree
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shortleaf pine (Pinus echinata), tree
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winged elm (Ulmus alata), tree
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white ash (Fraxinus americana), tree
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coralberry (Symphoricarpos orbiculatus), shrub
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blackhaw (Viburnum prunifolium), shrub
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rusty blackhaw (Viburnum rufidulum), shrub
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bursting-heart (Euonymus americanus), shrub
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eastern sweetshrub (Calycanthus floridus), shrub
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rosette grass (Dichanthelium), grass
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broad looseflower sedge (Carex laxiflora), grass
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rosy sedge (Carex rosea), grass
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eastern woodland sedge (Carex blanda), grass
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poverty oatgrass (Danthonia spicata), grass
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downy danthonia (Danthonia sericea), grass
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twoflower melicgrass (Melica mutica), grass
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nodding fescue (Festuca subverticillata), grass
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hairy woodland brome (Bromus pubescens), grass
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bearded shorthusk (Brachyelytrum erectum), grass
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eastern bottlebrush grass (Elymus hystrix), grass
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nakedflower ticktrefoil (Desmodium nudiflorum), other herbaceous
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feathery false lily of the valley (Maianthemum racemosum ssp. racemosum), other herbaceous
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ebony spleenwort (Asplenium platyneuron), other herbaceous
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dimpled troutlily (Erythronium umbilicatum), other herbaceous
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smooth Solomon's seal (Polygonatum biflorum), other herbaceous
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licorice bedstraw (Galium circaezans), other herbaceous
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Virginia snakeroot (Aristolochia serpentaria), other herbaceous
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wreath goldenrod (Solidago caesia), other herbaceous
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skullcap (Scutellaria), other herbaceous
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wild comfrey (Cynoglossum virginianum), other herbaceous
Community 1.2
Moist Basic Oak-Hickory Forest - Fire Suppressed PhaseThis is a closed canopy mature forest community/phase. This phase accounts for the majority of contemporary examples. Canopy cover is higher than in stands in which fire has been reintroduced. The pine component can have a greater proportion of loblolly or Virginia pine and the understory usually contains a greater proportion of fire-intolerant species. The herbaceous understory is typically sparse.
Forest overstory.The overstory is dominated by oaks. Representative species include white oak (Quercus alba) and northern red oak (Quercus rubra), with a smaller but significant contribution from hickories. Pines are typically scattered throughout the forest. Characteristic hickory species include shagbark hickory (Carya ovata), southern shagbark hickory (C. carolinae-septentrionalis), and red hickory (C. ovalis), along with the more widespread pignut and mockernut hickories (C. glabra, C. tomentosa). Loblolly pine (Pinus taeda) and shortleaf pine (Pinus echinata) are most representative of the pine component. In portions of the MLRA, Virginia pine (Pinus virginiana) can be equally important in fire suppressed stands.
Forest understory. Characteristic understory tree species include eastern redbud (Cercis canadensis), flowering dogwood (Cornus florida), white ash (Fraxinus americana), and hickory (Carya spp.), along with fire-intolerant species such as hophornbeam (Ostrya virginiana), southern sugar maple (Acer floridanum), red maple (Acer rubrum), American beech (Fagus grandifolia), and American holly (Ilex opaca).<br /> <br /> Characteristic understory shrub species include blackhaw (Viburnum prunifolium), bursting-heart (Euonymus americanus), American holly (Ilex opaca), rusty blackhaw (Viburnum rufidulum), and eastern sweetshrub (Calycanthus floridus), along with several vines.<br /> <br /> The herb layer is sparser and less diverse than in the fire maintained phase.
Dominant plant species
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white oak (Quercus alba), tree
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northern red oak (Quercus rubra), tree
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hybrid hickory (Carya), tree
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southern red oak (Quercus falcata), tree
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eastern redbud (Cercis canadensis), tree
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hophornbeam (Ostrya virginiana), tree
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flowering dogwood (Cornus florida), tree
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southern sugar maple (Acer floridanum), tree
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loblolly pine (Pinus taeda), tree
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red maple (Acer rubrum), tree
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American beech (Fagus grandifolia), tree
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blackhaw (Viburnum prunifolium), shrub
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bursting-heart (Euonymus americanus), shrub
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American holly (Ilex opaca), shrub
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muscadine (Vitis rotundifolia), shrub
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Virginia creeper (Parthenocissus quinquefolia), shrub
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rusty blackhaw (Viburnum rufidulum), shrub
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eastern sweetshrub (Calycanthus floridus), shrub
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nakedflower ticktrefoil (Desmodium nudiflorum), other herbaceous
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feathery false lily of the valley (Maianthemum racemosum ssp. racemosum), other herbaceous
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ebony spleenwort (Asplenium platyneuron), other herbaceous
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dimpled troutlily (Erythronium umbilicatum), other herbaceous
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licorice bedstraw (Galium circaezans), other herbaceous
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smooth Solomon's seal (Polygonatum biflorum), other herbaceous
Pathway 1.1A
Community 1.1 to 1.2Long-term exclusion of fire causes an increase in fire-intolerant understory species and a deterioration of the abundance and diversity of herbaceous species.
Key drivers
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Fire
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Lack of fire
Key ecosystem services affected
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Nutrient cycling
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Wildlife habitat
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Plant biodiversity
Pathway 1.2A
Community 1.2 to 1.1The fire suppressed phase can be managed towards the fire maintained phase through a combination of prescribed burns and selective removals. To approximate the pre-colonial fire regime, prescribed burns should be carried out every 4 to 8 years.
Context dependence.After decades of fire suppression, most upland hardwood forests of the Southeast have undergone mesophication, or succession toward forest systems that are less apt to burn. If prescribed fire is to be used as a management tool in fire suppressed ecosystems of the Piedmont, planning will be needed in some forest systems to overcome the effects of mesophication in the early stages of fire reintroduction.
Key drivers
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Fire
Key ecosystem services affected
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Nutrient cycling
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Wildlife habitat
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Plant biodiversity
State 2
Secondary Succession StateThis state develops in the immediate aftermath of agricultural abandonment, clearcut logging, or other large-scale disturbances that lead to canopy removal. Which species colonize a particular location in the wake of a disturbance does involve a considerable degree of chance. It also depends a great deal on the type, duration, and magnitude of the disturbance event.
Characteristics and indicators. Plant age distribution is even. Plants exhibit pioneering traits such as rapid growth, early reproduction, and shade-intolerance.
Community 2.1
Old-field Pine-Hardwood Forest PhaseThis forested successional phase develops in the wake of long-term agricultural abandonment or other large-scale disturbances that have led to canopy removal in the recent past. Stands are even-aged and species diversity is low. The canopy is usually dominated by pines, though opportunistic hardwoods can also be important, particularly in the early stages of tree establishment. Species that exhibit pioneering traits are usually most abundant.
Forest overstory.The overstory is dominated by pines. Loblolly pine (Pinus taeda) is the most characteristic species, followed by shortleaf pine (P. echinata), and to the north and west Virginia pine (P. virginiana).
Forest understory. Sweetgum (Liquidambar styraciflua) is the most representative understory tree species, though its importance tends to decline as the pines mature. Other common species include red maple (Acer rubrum), eastern redcedar (Juniperus virginiana), and American beech (Fagus grandifolia). Seedlings of oaks and hickories are usually present in the understory. These seedlings are released gradually as the forest matures and the pines begin to die off
Dominant plant species
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loblolly pine (Pinus taeda), tree
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sweetgum (Liquidambar styraciflua), tree
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red maple (Acer rubrum), tree
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eastern redcedar (Juniperus virginiana), tree
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Virginia pine (Pinus virginiana), tree
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shortleaf pine (Pinus echinata), tree
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American beech (Fagus grandifolia), tree
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tuliptree (Liriodendron tulipifera), tree
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black cherry (Prunus serotina), tree
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eastern redbud (Cercis canadensis), tree
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oak (Quercus), tree
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hybrid hickory (Carya), tree
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American holly (Ilex opaca), shrub
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muscadine (Vitis rotundifolia), shrub
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Virginia creeper (Parthenocissus quinquefolia), shrub
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Japanese honeysuckle (Lonicera japonica), shrub
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eastern poison ivy (Toxicodendron radicans), shrub
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Chinese privet (Ligustrum sinense), shrub
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autumn olive (Elaeagnus umbellata), shrub
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greenbrier (Smilax), shrub
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blueberry (Vaccinium), shrub
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trumpet creeper (Campsis radicans), shrub
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littlehead nutrush (Scleria oligantha), grass
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ebony spleenwort (Asplenium platyneuron), other herbaceous
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sparselobe grapefern (Botrychium biternatum), other herbaceous
Community 2.2
Shrub-dominated Successional PhaseThis successional phase is dominated by shrubs and vines, along with seedlings of opportunistic hardwoods and pines. It typically develops beginning in the third year after agricultural abandonment or clearcut logging. It grades into the forested successional phase as tree seedlings become saplings and begin to occupy more of the canopy cover.
Forest overstory.Species composition varies considerably from location to location. Exotic species usually occupy some portion of the vine or shrub cover in most examples.
Dominant plant species
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sweetgum (Liquidambar styraciflua), tree
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loblolly pine (Pinus taeda), tree
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Callery pear (Pyrus calleryana), tree
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Chinaberrytree (Melia azedarach), tree
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winged elm (Ulmus alata), tree
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eastern redcedar (Juniperus virginiana), tree
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black cherry (Prunus serotina), tree
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white ash (Fraxinus americana), tree
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silktree (Albizia julibrissin), tree
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honeylocust (Gleditsia triacanthos), tree
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tree of heaven (Ailanthus altissima), tree
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blackberry (Rubus), shrub
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rose (Rosa), shrub
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Japanese honeysuckle (Lonicera japonica), shrub
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winged sumac (Rhus copallinum), shrub
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autumn olive (Elaeagnus umbellata), shrub
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Chinese privet (Ligustrum sinense), shrub
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greenbrier (Smilax), shrub
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eastern poison ivy (Toxicodendron radicans), shrub
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coralberry (Symphoricarpos orbiculatus), shrub
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sweet autumn virginsbower (Clematis terniflora), shrub
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Johnsongrass (Sorghum halepense), grass
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broomsedge bluestem (Andropogon virginicus), grass
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Canada goldenrod (Solidago altissima), other herbaceous
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dogfennel (Eupatorium capillifolium), other herbaceous
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aster (Symphyotrichum), other herbaceous
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sericea lespedeza (Lespedeza cuneata), other herbaceous
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Indianhemp (Apocynum cannabinum), other herbaceous
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yellow crownbeard (Verbesina occidentalis), other herbaceous
Community 2.3
Herbaceous Early Successional PhaseThis transient community is composed of the first herbaceous invaders in the aftermath of agricultural abandonment, clearcut logging, or other large-scale natural disturbances that lead to canopy removal.
Species composition is highly variable at this stage of succession. In addition to the named species, other herbaceous pioneers common to this ecological site include wild lettuce (Lactuca spp.), fleabane (Erigeron spp.), Carolina horsenettle (Solanum carolinense), sericea lespedeza (Lespedeza cuneata), yellow crownbeard (Verbesina occidentalis), vetch (Vicia spp.), dock (Rumex spp.), dwarf dandelion (Krigia virginica), cudweed (Pseudognaphalium spp.), dwarf cinquefoil (Potentilla canadensis), Virginia threeseed mercury (Acalypha virginica), evening primrose (Oenothera spp.), great ragweed (Ambrosia trifida), starry rosinweed (Silphium asteriscus), several species of thoroughwort (Eupatorium spp.), and many others.
Resilience management. If the user wishes to maintain this community/phase for wildlife or pollinator habitat, a prescribed burn, mowing, or prescribed grazing will be needed at least once annually to prevent community pathway 2.3A. To that end, as part of long-term maintenance, periodic overseeding of wildlife or pollinator seed mixtures can be helpful in ensuring the viability of certain desired species and maintaining the desired composition of species for user goals.
Dominant plant species
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Japanese honeysuckle (Lonicera japonica), shrub
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greenbrier (Smilax), shrub
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broomsedge bluestem (Andropogon virginicus), grass
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hairy crabgrass (Digitaria sanguinalis), grass
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smooth crabgrass (Digitaria ischaemum), grass
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southern crabgrass (Digitaria ciliaris), grass
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Johnsongrass (Sorghum halepense), grass
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annual bluegrass (Poa annua), grass
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Canada goldenrod (Solidago altissima), other herbaceous
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American burnweed (Erechtites hieraciifolius), other herbaceous
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dogfennel (Eupatorium capillifolium), other herbaceous
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Canadian horseweed (Conyza canadensis), other herbaceous
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annual ragweed (Ambrosia artemisiifolia), other herbaceous
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American pokeweed (Phytolacca americana), other herbaceous
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aster (Symphyotrichum), other herbaceous
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Indianhemp (Apocynum cannabinum), other herbaceous
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Queen Anne's lace (Daucus carota), other herbaceous
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beggarticks (Bidens), other herbaceous
Pathway 2.1A
Community 2.1 to 2.3The old-field pine-hardwood forest phase can return to the herbaceous early successional phase through clearcut logging or other large-scale disturbances that cause canopy removal.
Context dependence.Note: if the user wishes to use this community pathway to create wildlife or pollinator habitat, please contact a local NRCS office for a species list specific to the area of interest and user needs.
Key drivers
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Timber management
Pathway 2.2A
Community 2.2 to 2.1The shrub-dominated successional phase naturally moves towards the old-field pine-hardwood forest through natural succession.
Pathway 2.2B
Community 2.2 to 2.3The shrub-dominated successional phase can return to the herbaceous early successional phase through brush management, including herbicide application, mechanical removal, prescribed grazing, or fire.
Context dependence.Note: if the user wishes to use this community pathway to create wildlife or pollinator habitat, please contact a local NRCS office for a species list specific to the area of interest and user needs. If the user wishes to maintain the shrub-dominated successional phase long term, for wildlife habitat or other uses, periodic use of this community pathway is necessary to prevent community pathway 2.2A, which happens inevitably unless natural succession is set back through disturbance.
Key drivers
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Fire
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Livestock grazing or browsing
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Brush management initiated by humans.
Pathway 2.3A
Community 2.3 to 2.2The herbaceous early successional phase naturally moves towards the shrub-dominated successional phase through natural succession. The process takes approximately 3 years on average, barring any major disturbances capable of inhibiting natural succession.
State 3
High-graded Hardwood Forest StateThis state develops as a consequence of high-grading, where the most valuable trees are removed, leaving less desirable timber specimens behind. Trees left behind include undesirable timber species, trees of poor form, diseased trees, or genetically inferior trees.
Characteristics and indicators. Typically, high-graded stands consist of a combination of residual stems from the previous stand, a high proportion of undesirable shade-tolerant species, along with some regrowth from desirable timber species. In some cases, large-diameter trees of desirable timber species may be present, but upon closer inspection, these trees usually have serious defects that resulted in their being left behind in earlier cuts.
Resilience management. Landowners with high-graded stands have two options for improving timber production: 1) rehabilitate, or 2) regenerate. To rehabilitate a stand, the landowner must evaluate existing trees to determine if rehabilitation is justified. If the proportion of high-quality specimens present in the stand is low, then the stand should be regenerated. In many cases, poor quality of the existing stand is the result of decades of mismanagement. Drastic measures are often required to get the stand back into good timber production.
Dominant plant species
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elm (Ulmus), tree
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maple (Acer), tree
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American beech (Fagus grandifolia), tree
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American holly (Ilex opaca), tree
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common persimmon (Diospyros virginiana), tree
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eastern redbud (Cercis canadensis), tree
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eastern redcedar (Juniperus virginiana), tree
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oak (Quercus), tree
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hybrid hickory (Carya), tree
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pine (Pinus), tree
State 4
Managed Pine Plantation StateThis converted state is dominated by planted timber trees. Loblolly pine (Pinus taeda) is the most commonly planted species. Even-aged management is the most common timber management system.
Note: if the user wishes to convert stands dominated by hardwoods to planted pine, clearcutting will usually be necessary first, allowing herbaceous pioneers to establish on the site in the weeks or months prior to planting. Users should utilize measures described in transition T2B under these circumstances.
Resilience management. Hardwood Encroachment: Hardwood encroachment can be problematic in managed pine plantations. Good site preparation, proper stocking, and periodic thinning are advisable to reduce hardwood competition. Overstocking: The overstocked condition commonly occurs in naturally regenerated stands. When competition from other pines begins to impact the health and productivity of the stand, precommercial thinning should be considered. At this point, the benefit of thinning usually outweighs the potential for invasion and competition from non-pine species. As the target window for thinning passes, the condition of the stand can slowly deteriorate if no action is taken. Under long-term overstocked conditions, trees are more prone to stresses, including pine bark beetle infestation and damage from wind or ice. High-grading: In subsequent commercial thinnings, care should be taken in tree selection. High quality specimens should be left to reach maturity, while slower growing trees or those with defects should be removed sooner. If high quality specimens are harvested first, trees left behind are often structurally unsound, diseased, genetically inferior, or of poor form. This can have long-term implications for tree genetics and for the condition of the stand (Felix III 1983; Miller et al. 1995, 2003; Megalos 2019).
Dominant plant species
-
loblolly pine (Pinus taeda), tree
-
sweetgum (Liquidambar styraciflua), tree
-
red maple (Acer rubrum), tree
-
eastern redcedar (Juniperus virginiana), tree
-
black cherry (Prunus serotina), tree
-
flowering dogwood (Cornus florida), tree
-
eastern redbud (Cercis canadensis), tree
-
oak (Quercus), tree
-
hybrid hickory (Carya), tree
-
American holly (Ilex opaca), shrub
-
grape (Vitis), shrub
-
Japanese honeysuckle (Lonicera japonica), shrub
-
greenbrier (Smilax), shrub
-
blackberry (Rubus), shrub
-
St. Johnswort (Hypericum), shrub
-
autumn olive (Elaeagnus umbellata), shrub
-
blueberry (Vaccinium), shrub
-
rosette grass (Dichanthelium), grass
-
longleaf woodoats (Chasmanthium sessiliflorum), grass
-
littlehead nutrush (Scleria oligantha), grass
-
silver plumegrass (Saccharum alopecuroides), grass
-
sortbeard plumegrass (Saccharum brevibarbe var. contortum), grass
-
broomsedge bluestem (Andropogon virginicus), grass
-
ebony spleenwort (Asplenium platyneuron), other herbaceous
-
sericea lespedeza (Lespedeza cuneata), other herbaceous
-
aster (Symphyotrichum), other herbaceous
-
thoroughwort (Eupatorium), other herbaceous
State 5
Pasture/Hayland StateThis converted state is dominated by herbaceous forage species.
Resilience management. Overgrazing and High Foot Traffic: In areas that are subject to high foot traffic from livestock and equipment, and/or long-term overgrazing, unpalatable weedy species tend to invade, as most desirable forage species are less competitive under these conditions. High risk areas include locations where livestock congregate for water, shade, or feed, and in travel lanes, gates, and other areas of heavy use. Plant species that are indicative of overgrazing or excessive foot traffic on this ecological site include buttercup (Ranunculus spp.), plantain (Plantago spp.), curly dock (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=RUCR" target="_blank" title="Open in plants.usda.gov"><i>Rumex crispus</i></a>), sneezeweed (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=HEAM" target="_blank" title="Open in plants.usda.gov"><i>Helenium amarum</i></a>), cudweed (Pseudognaphalium spp.), slender yellow woodsorrel (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=OXDI2" target="_blank" title="Open in plants.usda.gov"><i>Oxalis dillenii</i></a>), Carolina horsenettle (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=SOCA3" target="_blank" title="Open in plants.usda.gov"><i>Solanum carolinense</i></a>), Virginia pepperweed (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=LEVI3" target="_blank" title="Open in plants.usda.gov"><i>Lepidium virginicum</i></a>), black medick (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=MELU" target="_blank" title="Open in plants.usda.gov"><i>Medicago lupulina</i></a>), Japanese clover (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=KUST2" target="_blank" title="Open in plants.usda.gov"><i>Kummerowia striata</i></a>), annual bluegrass (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=POAN" target="_blank" title="Open in plants.usda.gov"><i>Poa annua</i></a>), poverty rush (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=JUTE" target="_blank" title="Open in plants.usda.gov"><i>Juncus tenuis</i></a>), rattail fescue (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=VUMY" target="_blank" title="Open in plants.usda.gov"><i>Vulpia myuros</i></a>), and Indian goosegrass (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=ELIN3" target="_blank" title="Open in plants.usda.gov"><i>Eleusine indica</i></a>), among others. A handful of desirable forage species are also tolerant of heavy grazing and high foot traffic, including white clover (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=TRRE3" target="_blank" title="Open in plants.usda.gov"><i>Trifolium repens</i></a>), dallisgrass (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=PADI3" target="_blank" title="Open in plants.usda.gov"><i>Paspalum dilatatum</i></a>), and bermudagrass (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=CYDA" target="_blank" title="Open in plants.usda.gov"><i>Cynodon dactylon</i></a>). An overabundance of these species, along with poor plant vigor and areas of bare soil, may imply that excessive foot traffic and/or overgrazing is a concern, either in the present or in the recent past. Brush Encroachment: Brush encroachment can be problematic in some pastures, particularly near fence lines where there is often a ready seed source. Pastures subject to low stocking density and long-duration grazing rotations can also be susceptible to encroachment from woody plants. Shorter grazing rotations of higher stocking density can help alleviate pressure from shrubs and vines with low palatability or thorny stems. Clipping behind grazing rotations, annual brush hogging, and multispecies grazing systems (cattle with or followed by goats) can also be helpful. Common woody invaders of pasture on this ecological site include rose (Rosa spp.), blackberry (Rubus spp.), saw greenbrier (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=SMBO2" target="_blank" title="Open in plants.usda.gov"><i>Smilax bona-nox</i></a>), Japanese honeysuckle (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=LOJA" target="_blank" title="Open in plants.usda.gov"><i>Lonicera japonica</i></a>), common persimmon (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=DIVI5" target="_blank" title="Open in plants.usda.gov"><i>Diospyros virginiana</i></a>), eastern redcedar (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=JUVI" target="_blank" title="Open in plants.usda.gov"><i>Juniperus virginiana</i></a>), black cherry (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=PRSE2" target="_blank" title="Open in plants.usda.gov"><i>Prunus serotina</i></a>), and Chinese privet (<a class="species-link" href="https://plants.usda.gov/core/profile?symbol=LISI" target="_blank" title="Open in plants.usda.gov"><i>Ligustrum sinense</i></a>).
Dominant plant species
-
tall fescue (Schedonorus arundinaceus), grass
-
Bermudagrass (Cynodon dactylon), grass
-
orchardgrass (Dactylis glomerata), grass
-
dallisgrass (Paspalum dilatatum), grass
-
perennial ryegrass (Lolium perenne), grass
-
Johnsongrass (Sorghum halepense), grass
-
hairy crabgrass (Digitaria sanguinalis), grass
-
purpletop tridens (Tridens flavus), grass
-
bahiagrass (Paspalum notatum), grass
-
broomsedge bluestem (Andropogon virginicus), grass
-
white clover (Trifolium repens), other herbaceous
-
red clover (Trifolium pratense), other herbaceous
-
vetch (Vicia), other herbaceous
-
narrowleaf plantain (Plantago lanceolata), other herbaceous
-
dogfennel (Eupatorium capillifolium), other herbaceous
-
black medick (Medicago lupulina), other herbaceous
-
field clover (Trifolium campestre), other herbaceous
-
common dandelion (Taraxacum officinale), other herbaceous
-
wild garlic (Allium vineale), other herbaceous
State 6
Cropland StateThis converted state produces food or fiber for human uses. It is dominated by domesticated crop species, along with typical weedy invaders of cropland.
Community 6.1
Conservation-management Cropland PhaseThis cropland phase is characterized by the practice of no-tillage or strip-tillage, and other soil conservation practices. Though no-till systems offer many benefits, several weedy species tend to be more problematic under this type of management system. In contrast with conventional tillage systems, problematic species in no-till systems include biennial or perennial weeds, owing to the fact that tillage is no longer used in weed management.
Dominant plant species
-
corn (Zea mays), grass
-
common wheat (Triticum aestivum), grass
-
grain sorghum (Sorghum bicolor ssp. bicolor), grass
-
soybean (Glycine max), other herbaceous
-
upland cotton (Gossypium hirsutum), other herbaceous
-
cultivated tobacco (Nicotiana tabacum), other herbaceous
Community 6.2
Conventional-management Cropland PhaseThis cropland phase is characterized by the recurrent use of tillage as a management tool. Due to the frequent disturbance regime, weedy invaders tend to be annual herbaceous species that reproduce quickly and are prolific seed producers.
Resilience management. The potential for soil loss is high under this management system. Measures should be put in place to limit erosion.
Dominant plant species
-
corn (Zea mays), grass
-
common wheat (Triticum aestivum), grass
-
grain sorghum (Sorghum bicolor ssp. bicolor), grass
-
soybean (Glycine max), other herbaceous
-
upland cotton (Gossypium hirsutum), other herbaceous
-
cultivated tobacco (Nicotiana tabacum), other herbaceous
Pathway 6.1A
Community 6.1 to 6.2The conservation-management cropland phase can shift to the conventional-management cropland phase through cessation of conservation tillage practices and the reintroduction of conventional tillage practices.
Context dependence.Soil and vegetation changes associated with this community pathway typically occur several years after reintroduction of conventional tillage practices. These changes continue to manifest as conventional tillage is continued, before reaching a steady state.
Key drivers
-
Mechanical soil disturbance
Key ecosystem services affected
-
Erosion control
-
Climate regulation: carbon cycling and storage
-
Food and fiber: crops
Pathway 6.2A
Community 6.2 to 6.1The conventional-management cropland phase can be brought into the conservation-management cropland phase through the implementation of one of several conservation tillage options, including no-tillage or strip-tillage, along with implementation of other soil conservation practices.
Context dependence.Soil and vegetation changes associated with this community pathway typically occur several years after implementation of conservation tillage. These changes continue to manifest as conservation tillage is continued, before reaching a steady state.
Key drivers
-
Mechanical soil disturbance
Key ecosystem services affected
-
Erosion control
-
Climate regulation: carbon cycling and storage
-
Food and fiber: crops
Transition T1A
State 1 to 2The reference state can transition to the secondary succession state through clearcut logging or other large-scale disturbances that cause canopy removal.
Key drivers
-
Timber management
Transition T1B
State 1 to 3The reference state can transition to the high-graded hardwood forest state through selective removal of the most valuable trees, leaving undesirable timber specimens behind. This may occur through multiple cutting cycles over the course of decades or longer, each cut progressively worsening the condition of the stand.
Key drivers
-
Timber management
Key ecosystem services affected
-
Food and fiber: wood products
-
Plant biodiversity
Transition T1C
State 1 to 5The reference state can transition to the pasture/hayland state through 1) mechanical tree/brush/stump/debris removal, 2) seedbed preparation, and 3) planting of perennial grasses and forbs.
Context dependence.Herbicide applications, fire, and/or root-raking can be helpful in transitioning treed land to pasture. This is done in part to limit coppicing, as many woody plants are capable of sprouting from residual plant structures left behind after clearing. Judicious use of root-raking is recommended, as this practice can have long-term repercussions with regard to soil structure. Applications of fertilizer and lime can also be helpful in establishing perennial forage species. Grazing should be deferred until grasses and forbs are well established.
Key drivers
-
Mechanical soil disturbance
-
Timber management
-
Seeding
Key ecosystem services affected
-
Cultural heritage values
-
Food and fiber: livestock forage
-
Plant biodiversity
-
Natural heritage values
Transition T1D
State 1 to 6The reference state can transition to the cropland state through 1) mechanical tree/brush/stump/debris removal, 2) seedbed preparation, 3) applications of fertilizer/lime, and 4) planting of crop or cover crop seed.
Context dependence.A broad spectrum herbicide, fire, and/or root-raking can be helpful in transitioning treed land to cropland. This is done in part to limit coppicing, as many woody plants are capable of sprouting from residual plant structures left behind after clearing. Judicious use of root-raking is recommended, as this practice can have long-term repercussions with regard to soil structure. Weedy grasses and forbs can also be problematic on these lands.
Key drivers
-
Mechanical soil disturbance
-
Timber management
-
Seeding
Key ecosystem services affected
-
Cultural heritage values
-
Erosion control
-
Climate regulation: carbon cycling and storage
-
Wildlife habitat
-
Plant biodiversity
-
Natural heritage values
-
Food and fiber: crops
Transition T2A
State 2 to 1The secondary succession state can transition to the reference state through long-term natural succession. This process can be accelerated to some degree by a combination of prescribed burns and selective harvesting of pines and opportunistic hardwoods.
Transition T2B
State 2 to 4The secondary succession state can transition to the managed pine plantation state through site preparation and planting of timber trees. Thinning alone may be sufficient for portions of the forest if pines have already established, though it is rarely sufficient for an entire forest patch.
Key drivers
-
Timber management
Key ecosystem services affected
-
Food and fiber: wood products
Transition T2C
State 2 to 5The secondary succession state can transition to the pasture/hayland state through through 1) mechanical tree/brush/stump/debris removal, 2) seedbed preparation, and 3) planting of perennial grasses and forbs.
Context dependence.A broad spectrum herbicide, fire, and/or root-raking can be helpful in transitioning wooded or semi-wooded land to pasture. This is done in part to limit coppicing, as many woody pioneers are capable of sprouting from residual plant structures left behind after clearing. Judicious use of root-raking is recommended, as this practice can have long-term repercussions with regard to soil structure. Applications of fertilizer and lime can also be helpful in establishing perennial forage species. Grazing should be deferred until grasses and forbs are well established.
Key drivers
-
Livestock grazing or browsing
-
Seeding
-
Brush management
Key ecosystem services affected
-
Food and fiber: livestock forage
Transition T2D
State 2 to 6The secondary succession state can transition to the cropland state through 1) mechanical tree/brush/stump/debris removal, 2) seedbed preparation, 3) applications of fertilizer/lime, 4) weed control, 5) planting of crop or cover crop seed.
Context dependence.A broad spectrum herbicide, fire, and/or root-raking may be needed to successfully transition land that has been fallow for some time back to cropland. This is done in part to limit coppicing, as many woody pioneers are capable of sprouting from residual plant structures left behind after clearing. Judicious use of root-raking is recommended, as this practice can have long-term repercussions with regard to soil structure. Weedy grasses and forbs can also be problematic on these lands.
Key drivers
-
Mechanical soil disturbance
-
Seeding
-
Mechanical land clearing
Key ecosystem services affected
-
Erosion control
-
Climate regulation: carbon cycling and storage
Transition T3A
State 3 to 2The high-graded hardwood forest state can transition to the secondary succession state through clearcut logging or other large-scale disturbances that cause canopy removal.
Key drivers
-
Timber management
Transition T3C
State 3 to 5The high-graded hardwood forest state can transition to the pasture/hayland state through 1) mechanical tree/brush/stump/debris removal, 2) seedbed preparation, and 3) planting of perennial grasses and forbs.
Context dependence.Herbicide applications, fire, and/or root-raking can be helpful in transitioning treed land to pasture. This is done in part to limit coppicing, as many woody plants are capable of sprouting from residual plant structures left behind after clearing. Judicious use of root-raking is recommended, as this practice can have long-term repercussions with regard to soil structure. Applications of fertilizer and lime can also be helpful in establishing perennial forage species. Grazing should be deferred until grasses and forbs are well established.
Key drivers
-
Mechanical soil disturbance
-
Timber management
-
Seeding
Key ecosystem services affected
-
Food and fiber: livestock forage
Transition T3D
State 3 to 6The high-graded hardwood forest state can transition to the cropland state through 1) mechanical tree/brush/stump/debris removal, 2) seedbed preparation, 3) applications of fertilizer/lime, 4) weed control, 5) planting of crop or cover crop seed.
Constraints to recovery.A broad spectrum herbicide, fire, and/or root-raking can be helpful in transitioning treed land to cropland. This is done in part to limit coppicing, as many woody pioneers are capable of sprouting from residual plant structures left behind after clearing. Judicious use of root-raking is recommended, as this practice can have long-term repercussions with regard to soil structure. Weedy grasses and forbs can also be problematic on these lands.
Key drivers
-
Mechanical soil disturbance
-
Timber management
-
Seeding
Key ecosystem services affected
-
Erosion control
-
Climate regulation: carbon cycling and storage
-
Plant biodiversity
-
Food and fiber: crops
Transition T4A
State 4 to 2The managed pine plantation state can transition to the secondary succession state through abandonment of forestry practices (with or without timber tree harvest).
Key drivers
-
Timber management
Key ecosystem services affected
-
Food and fiber: wood products
Transition T4B
State 4 to 5The managed pine plantation state can transition to the pasture/hayland state through 1) timber harvest, 2) mechanical stump and debris removal, 3) seedbed preparation, 4) planting of perennial grasses and forbs.
Context dependence.Applications of fertilizer and lime can be helpful in establishing perennial forage species. Grazing should be deferred until grasses and forbs are well established.
Key drivers
-
Mechanical soil disturbance
-
Timber management
-
Seeding
Key ecosystem services affected
-
Food and fiber: livestock forage
-
Food and fiber: wood products
Transition T4C
State 4 to 6The managed pine plantation state can transition to the cropland state through 1) timber harvest, 2) mechanical stump and debris removal, 3) seedbed preparation, 4) applications of fertilizer/lime, 5) weed control, 6) planting of crop or cover crop seed.
Key drivers
-
Mechanical soil disturbance
-
Timber management
-
Seeding
Key ecosystem services affected
-
Erosion control
-
Food and fiber: wood products
-
Food and fiber: crops
Transition T5A
State 5 to 2The pasture/hayland state can transition to the secondary succession state through long-term cessation of grazing.
Key ecosystem services affected
-
Aesthetic values
-
Food and fiber: livestock forage
Transition T5B
State 5 to 4The pasture/hayland state can transition to the managed pine plantation state through site preparation and tree planting.
Key drivers
-
Timber management
Key ecosystem services affected
-
Food and fiber: livestock forage
-
Food and fiber: wood products
Transition T5C
State 5 to 6The pasture/hayland state can transition to the cropland state through 1) seedbed preparation, 2) applications of fertilizer/lime, 3) weed control, and 4) planting of crop or cover crop seed.
Key drivers
-
Mechanical soil disturbance
-
Seeding
Key ecosystem services affected
-
Erosion control
-
Food and fiber: livestock forage
-
Climate regulation: carbon cycling and storage
-
Food and fiber: crops
Transition T6A
State 6 to 2The cropland state can transition to the secondary succession state through agricultural abandonment.
Transition T6B
State 6 to 4The cropland state can transition to the managed pine plantation state through site preparation and tree planting.
Key drivers
-
Timber management
-
Food and fiber: crops
Key ecosystem services affected
-
Food and fiber: wood products
-
Food and fiber: crops
Transition T6C
State 6 to 5The cropland state can transition to the pasture/hayland state through 1) seedbed preparation, 2) weed control, and 3) planting of perennial forage grasses and forbs.
Context dependence.To convert cropland to pasture or hayland, weed control and good seed-soil contact are important. It is also critical to review the labels of herbicides used for weed control and on the previous crop. Many herbicides have plant-back restrictions, which if not followed could carryover and kill forage seedlings as they germinate. Grazing should be deferred until grasses and forbs are well established.
Key drivers
-
Mechanical soil disturbance
-
Seeding
Key ecosystem services affected
-
Erosion control
-
Food and fiber: livestock forage
-
Climate regulation: carbon cycling and storage
-
Food and fiber: crops
Additional community tables
Table 10. Community 1.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 11. Community 1.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 12. Community 2.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 13. Community 2.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 14. Community 2.3 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 15. Community 6.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 16. Community 6.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Interpretations
Supporting information
Inventory data references
Data collection and analysis of field data will be performed during the Verification Stage of ESD development.
Other references
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Yogev Erez
Dee PedersonApproval
Charles Stemmans, 5/02/2025
Rangeland health reference sheet
Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.
Author(s)/participant(s) Contact for lead author Date 07/14/2026 Approved by Approval date Composition (Indicators 10 and 12) based on Annual Production Indicators
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Number and extent of rills:
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Presence of water flow patterns:
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Number and height of erosional pedestals or terracettes:
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Bare ground from Ecological Site Description or other studies (rock, litter, lichen, moss, plant canopy are not bare ground):
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Number of gullies and erosion associated with gullies:
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Extent of wind scoured, blowouts and/or depositional areas:
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Amount of litter movement (describe size and distance expected to travel):
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Soil surface (top few mm) resistance to erosion (stability values are averages - most sites will show a range of values):
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Soil surface structure and SOM content (include type of structure and A-horizon color and thickness):
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Effect of community phase composition (relative proportion of different functional groups) and spatial distribution on infiltration and runoff:
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Presence and thickness of compaction layer (usually none; describe soil profile features which may be mistaken for compaction on this site):
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Functional/Structural Groups (list in order of descending dominance by above-ground annual-production or live foliar cover using symbols: >>, >, = to indicate much greater than, greater than, and equal to):
Dominant:
Sub-dominant:
Other:
Additional:
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Amount of plant mortality and decadence (include which functional groups are expected to show mortality or decadence):
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Average percent litter cover (%) and depth ( in):
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Expected annual annual-production (this is TOTAL above-ground annual-production, not just forage annual-production):
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Potential invasive (including noxious) species (native and non-native). List species which BOTH characterize degraded states and have the potential to become a dominant or co-dominant species on the ecological site if their future establishment and growth is not actively controlled by management interventions. Species that become dominant for only one to several years (e.g., short-term response to drought or wildfire) are not invasive plants. Note that unlike other indicators, we are describing what is NOT expected in the reference state for the ecological site:
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Perennial plant reproductive capability:
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PrintThe Ecosystem Dynamics Interpretive Tool is an information system framework developed by the USDA-ARS Jornada Experimental Range, USDA Natural Resources Conservation Service, and New Mexico State University.
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