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Ecological site F134XY208AL
Western Dry Loess Backslope
Last updated: 3/24/2025
Accessed: 08/06/2026
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Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.
MLRA notes
Major Land Resource Area (MLRA): 134X–Southern Mississippi Valley Loess
The Southern Mississippi Valley Loess (MLRA 134) extends some 500 miles from the southern tip of Illinois to southern Louisiana. This MLRA occurs in Mississippi (39 percent), Tennessee (23 percent), Louisiana (15 percent), Arkansas (11 percent), Kentucky (9 percent), Missouri (2 percent), and Illinois (1 percent). It makes up about 26,520 square miles. Landscapes consist of highly dissected uplands, level to undulating plains, and broad terraces that are covered with a mantle of loess. Underlying the loess are Tertiary deposits of unconsolidated sand, silt, clay, gravel, and lignite. The soils, mainly Alfisols, formed in the loess mantle. Stream systems of the MLRA typically originate as low-gradient drainageways in the upper reaches that broaden rapidly downstream to wide, level floodplains with highly meandering channels. Alluvial soils, mostly Entisols and Inceptisols, are predominantly silty where loess thickness of the uplands are deepest but grade to loamy textures in watersheds covered by thin loess. Crowley’s Ridge, Macon Ridge, and Lafayette Loess Plains are discontinuous, erosional remnants that run north to south in southeastern Missouri - eastern Arkansas, northeastern Louisiana, and south-central Louisiana, respectively. Elevations range from around 100 feet on terraces in southern Louisiana to over 600 feet on uplands in western Kentucky. The steep, dissected uplands are mainly in hardwood forests while less sloping areas are used for crop, pasture, and forage production (USDA-NRCS, 2006).
This site is restricted to the dry to moderately moist (i.e., mesic) backslopes of the northern section of Crowley’s Ridge from about Harrisburg in Poinsett County, Arkansas northward through portions of Stoddard County, Missouri.Classification relationships
All or portions of the geographic range of this site falls within a number of ecological/land classifications including:
-NRCS Major Land Resource Area (MLRA) 134 – Southern Mississippi Valley Loess
-Environmental Protection Agency’s Level IV Ecoregion: Bluff Hills, 74a (Griffith et al., 1998; Woods et al., 2002; Chapman et al., 2002; Chapman et al., 2004; Woods et al., 2004; Daigle et al., 2006)
-231H - Coastal Plains-Loess section of the USDA Forest Service Ecological Subregion (McNab et al., 2005)
-LANDFIRE Biophysical Setting 4515100 and NatureServe Ecological System CES203.072 Northern Crowley’s Ridge Sand Forest and Crowley’s Ridge Sand Forest, respectively (LANDFIRE, 2008; NatureServe, 2011)
-Dry Sand Woodland; Dry-Mesic Sand Woodland; Dry-Mesic Loess Forest; Dry-Mesic Loess Woodland; Dry-Mesic Sand Forest (Nelson, 2005)
-Western Mesophytic Forest Region - Mississippi Embayment Section - Loess Hills (Braun, 1950)
Ecological site concept
The Western Dry Loess Backslope site is restricted to the moderately steep to steep slopes of the northern sections of Crowley’s Ridge. Slopes of this site are greater than 12 percent and are frequently within the range of 12 to 35 percent. Soils of this site are deep and largely comprised of loess and loess over fluviomarine deposits (sand and gravel). Based on the distribution of soils, loess depths vary considerably with deep loess soils (greater than or equal to 4 feet thick), thin loess soils (greater than 20 inches and less than 40 inches thick), and exposed sand and gravel co-occurring in close proximity and within intricate complexes. Upper to mid-slope positions are tend to be quite droughty and this influence is reflected in the local plant community, which is often comprised of drier oaks, hickory, and shortleaf pine, where present. Lower slope positions (i.e., footslopes) often support species indicative of increased moisture such as beech, maple, tuliptree, sweetgum, and occasionally basswood and cucumber tree. This community is largely restricted to moist footslopes within narrow ravines and rarely, if ever, occurs beyond these protected environments. The driest community of this system occurs on local exposures of gravel and sand and on exposed slopes (south- to west-facing slopes). Here, shortleaf pine is often the dominant species with associates of post oak, blackjack oak, black hickory, and a heavy understory component of farkleberry. The contrasting plant communities occurring across this pronounced dry – moist gradient are treated and referenced in this context as dry – mesic forests (the usage of “mesic” in this case refers to moderate moisture and not soil temperature regime).
Associated sites
F134XY210AL Western Dry Loess Summit
Similar sites
F134XY006AL Northern Loess Sideslope
This site has similar soils and occurs on similar landforms as the Western Dry Loess Backslope site.
Table 1. Dominant plant species
Tree Not specified
Shrub Not specified
Herbaceous Not specified
Physiographic features
The Western Dry Loess Backslope ecological site occurs entirely within a distinct physiographic subsection of the Southern Mississippi Valley Loess (MLRA 134): Crowley’s Ridge. This prominent physiographic feature is a western counterpart to the Loess Hills east of the Mississippi River (Braun, 1950). Crowley’s Ridge is a narrow belt of low, dissected hills that extends roughly 200 miles north to south from southeastern Missouri into eastern Arkansas. Shared characteristics with the bluffs to the east include a loess-cap (but with varying depths) that is underlain by Tertiary deposits of silt, sand, clay, and gravel. One notable distinction of its geographic location is that the entire length of Crowley’s Ridge is surrounded by the Southern Mississippi River Alluvium (i.e., MLRA 131A; USDA, 2006) and is separated from the Loess Hills to the east by 23 to 50 miles of the vast Mississippi River delta region. Elevation crests over 500 feet above sea level with local topographic relief rising 200 feet above the adjoining alluvial plain (Clark et al., 1974). EPA combines Crowley’s Ridge and the bluffs to the east within a single Level IV Ecoregion: the Bluff Hills, 74a (Woods et al., 2004).
Although similarities exist between Crowley’s Ridge and the Loess Hills to the east (see Braun, 1950), there are some profound differences with respect to loess thickness. Loess depths thin markedly and is even absent in some areas through the northern section of the Ridge. An intricate complex of deep loess, thin loess, and exposed fluviomarine deposits is distributed across much of the area, and these physical differences have a direct influence on vegetation types.
This site occurs on moderately steep to steep backslopes and includes dry, exposed slopes and moist protected slopes.Table 2. Representative physiographic features
Landforms (1) Hill
(2) Ravine
Flooding frequency None Ponding frequency None Elevation 300 – 580 ft Slope 12 – 40 % Ponding depth 0 in Water table depth 60 in Aspect N, S, W Climatic features
This site falls under the Humid Subtropical Climate Classification (Koppen System). The mean annual precipitation for this site from 1980 through 2010 was approximately 49 inches with a range from 36 to roughly 65 inches. Maximum precipitation occurs in spring (April and May) and late fall (November and December) and typically decreases throughout the summer. Rainfall often occurs as high-intensity, convective thunderstorms during warmer periods but moderate-intensity frontal systems can produce large amounts of rainfall during winter. Snowfall generally occurs in most years, and the average annual snowfall in the northern portions of this site in Stoddard County, Missouri is 11 inches (USDA-NRCS, 2006). The average annual maximum and minimum air temperature is 69 and 48 degrees F, respectively. The average frost-free and freeze-free periods are 196 and 225 days, respectively.
Table 3 Representative climatic features
Frost-free period (average) 200 days Freeze-free period (average) 230 days Precipitation total (average) 50 in BarLineFigure 1. Monthly precipitation range
BarLineFigure 2. Monthly average minimum and maximum temperature
Figure 3. Annual precipitation pattern
Figure 4 Annual average temperature pattern
Climate stations used
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(1) PARAGOULD 1S [USC00035563], Paragould, AR
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(2) WYNNE [USC00038052], Wynne, AR
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(3) ADVANCE 1 S [USW00093825], Advance, MO
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(4) JONESBORO 2 NE [USC00033734], Jonesboro, AR
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(5) MALDEN MUNI AP [USC00235207], Malden, MO
">Influencing water features
This site is not influence by a hydrologic regime.
Soil features
The soils of this site are well drained and consist of a complex mixture of deep loess, thin loess, and exposed fluviomarine deposits (gravel and sand). A commonality among these soils is the dry association of plants that is supported; the exposed gravelly/sandy soils produce the driest and least productive plant community.
The deep loess soil (> 4 ft.) that has been mapped and is associated with this site is the Memphis series (Fine-silty, mixed, active, thermic Typic Hapludalfs). However, the dry plant association occurring on this site is unlike that observed on Memphis soils elsewhere in the MLRA. Base saturations (a measure of a soil’s natural fertility) for the Memphis series, a taxonomic criterion, should exceed 60 percent. It is unknown if the Memphis soils of this site meet that criterion.
Occurring within close proximity to many of the Memphis soil map units are Brandon soils and a Brandon – Saffell soil complex. Brandon soils (Fine-silty, mixed, semiactive, thermic Typic Hapludults) are the thin loess (< 4 ft.) counterpart to the Memphis soils. Brandon series consists of a thin loess mantle that is 20 to 40 inches thick over very gravelly or gravelly marine and riverine deposited materials. Solum thickness ranges from 20 to more than 48 inches. Rock fragments range from 0 to 5 percent in the solum and from 30 to 80 percent in the 2 Bt and 2C horizons. Reaction ranges from strongly to very strongly acid.
Saffell soils (Loamy-skeletal, siliceous, semiactive, thermic Typic Hapludults) formed in loamy and gravelly marine sediments of Tertiary Age. Soil reactions range from strongly acid to very strongly acid (USDA-NRCS, 2016).
The properties among the three soils that are associated with this site differ markedly. Their inclusion in this provisional site should be viewed as temporary (one of convenience) until future rigorous soil – site investigations better describes their differences (and/or similarities) with respect to plant community response and management. Final analysis may suggest that each soil component warrants its own separate and distinct ecological site.Table 4. Representative soil features
Surface texture (1) Gravelly silt loam
(2) Very gravelly loam
(3) Fine sandy loam
Family particle size (1) Loamy
Drainage class Well drained Permeability class Moderate to rapid Soil depth 80 in Surface fragment cover <=3" Not specified Surface fragment cover >3" Not specified Available water capacity
(0-40in)3.3 – 8.5 in Electrical conductivity
(0-40in)0 – 1 mmhos/cm Sodium adsorption ratio
(0-40in)Not specified Soil reaction (1:1 water)
(0-40in)5 – 5.6 Subsurface fragment volume <=3"
(Depth not specified)2 – 37 % Subsurface fragment volume >3"
(Depth not specified)2 – 5 % Ecological dynamics
This ecological site occurs on moderately steep to steep backslopes throughout portions of the northern section of Crowley’s Ridge. Landscape position coupled with well drained, droughty soils directly influence the plant types and productivity of the area. Unlike the prevailing soils to the south, which are mainly deep loess, a significant portion of the area consists of a complex distribution of thick loess (> 4 feet), thin loess (< 4 feet), and loamy-skeletal material that is mainly comprised of surface sand and gravel. This complexity of the physical environment is clearly expressed in plant community differences.
Reconnaissance of this site suggests that areas where a loess cap is thickest, a mixed oak – hickory association is often supported. Canopy components may vary by stand but in general, species include southern red oak, black oak, post oak, with white oak and northern red oak occurring on moist sites, especially protected aspects (i.e., northwest- to east-facing slopes). Hickories typically consist of shagbark, mockernut, pignut, and black, the latter occurring on drier sites. Where loess deposits are thin, the community shifts to a drier association with a greater concentration of post oak, black hickory, and an entrance or increased presence of shortleaf pine and blackjack oak. This association is most pronounced and developed on exposed aspects (i.e., southeast- to west-facing slopes). The relative abundance or dominance of community components may vary by stand with some sites dominated by oaks and yet others by shortleaf pine. Land use history of local sites likely have an important influence on composition of many stands. Within this complex mosaic of soils, the lower slopes (e.g., footslopes) consist of deep colluvium – the results of past erosion. The protected, moist conditions of the footslopes often support a higher concentration of beech, maple, tuliptree, basswood, and walnut.
The pre-settlement vegetation of Crowley’s Ridge was projected by Clark et al. (1974), which was drawn from the journals of early naturalists and published state geologic reports (e.g., Call, 1891). For the portion of the Ridge associated with this site, much of the area was classed as an Oak – Hickory – Pine forest with potential overlap with a predominantly Mixed Oak – Hickory type. Today, vestiges of these vegetation types occur in scattered locations. One likely difference between the pre-settlement communities and current conditions is reflected in structural characteristics. The pre-settlement plant communities on the drier slopes were very likely open woodlands where fire was a critical and recurring disturbance factor. Many fires likely extended onto the upper portions of the moist, protected slopes. However, those areas likely remained somewhat protected given their occurrence in “fire shadows” (Nelson, 2005) and may have functioned as natural fire breaks. The moist lower slopes likely persisted under forest conditions (i.e., canopy closure greater than 75 percent), artifacts of a fire-sheltered environment.
Once settlement commenced, land uses and habitat alteration were extensive. Any location that could grow crops was farmed. Much of the area was either converted to pasture or remained in timber and repeatedly logged. With the advent of fire suppression, regrowth of formerly cutover land was often overcrowded and productivity suppressed. Today, the major land uses continue to be pastureland and forestry.
Following this narrative, a “provisional” state and transition model is provided that includes the “perceived” reference state and several alternative (or altered) vegetation states that have been observed and/or projected for this ecological site. This model is based on limited inventories, literature, expert knowledge, and interpretations. Plant communities will differ across MLRA 134 due to natural variability in climate, soils, and physiography. Depending on objectives, the reference plant community may not necessarily be the management goal.
The environmental and biological characteristics of this site are complex and dynamic. As such, the following diagram suggests pathways that the vegetation on this site might take, given that the modal concepts of climate and soils are met within an area of interest. Specific locations with unique soils and disturbance histories may have alternate pathways that are not represented in the model. This information is intended to show the possibilities within a given set of circumstances and represents the initial steps toward developing a defensible description and model. The model and associated information are subject to change as knowledge increases and new information is garnered. This is an iterative process. Most importantly, local and/or state professional guidance should always be sought before pursuing a treatment scenario.State and transition model
Custom diagramStandard diagram
Figure 5. STM - Western Dry Loess Backslope
Figure 6. Legend - Western Dry Loess Backslope
More interactive model formats are also available. View Interactive Models
More interactive model formats are also available. View Interactive Models
Click on state and transition labels to scroll to the respective textEcosystem states
States 1, 5 and 2 (additional transitions)
State 1 submodel, plant communities
State 2 submodel, plant communities
State 3 submodel, plant communities
State 4 submodel, plant communities
State 5 submodel, plant communities
State 1
Old Growth Woodland and ForestThe pre-settlement plant community of this ecological site was largely removed more than 150 years ago, and there are no extant examples remaining. However, inferences over the structure and dynamics of that system are drawn based on landscape position, soils, and existing community components.
This site is distributed across a highly dissected terrain and accordingly, encompasses many complexities including the presence/absence of loess; varying loess depths, where present; exposed vs. protected aspects; and a pronounced moisture gradient from upper slopes to footslope positions. Reference conditions for this site vary naturally with respect to those physical differences and includes several distinct natural communities or associations.
Two community phases are recognized for the reference state and they are distinguished from one another mainly by succession and disturbance type, size, and frequency.
Community 1.1
Dry-Mesic Mixed Oak – Hickory – Shortleaf Pine Woodland and ForestThe dominant species that occur over the distribution of this site are highly varied and directly influenced by soil type, aspect, and landscape position. A large proportion of the site was likely influenced by periodic fire. As a whole, the system may be classed as fire-adapted (NatureServe, 2011). However, certain positions and locations such as moist footslopes and north-facing slopes may have developed full, overlapping canopies and functioned as natural fire breaks. This community phase is so named to include the broad compositional and structural variation over the distribution of this site.
Mid- to upper slope positions on exposed aspects likely supported woodland characteristics. Periodic fires helped to maintain an open to moderately open canopy and understory. On the driest sites, shortleaf pine was very likely the dominant species with associates consisting of post oak, southern red oak, blackjack oak, black oak, black hickory and an understory of lowbush blueberry, farkleberry, and aromatic sumac. Mid- to upper slopes on protected aspects likely supported an oak – hickory association that consisted of white oak, southern red oak, northern red oak, black oak, various hickories, black gum, and hophornbeam and dogwood as mid-story components. The lower slopes supported the largest trees and highest canopy coverage. This protected environment supported greater diversity of canopy components including American beech, tuliptree, sweetgum, basswood, maple, walnut, white oak, northern red oak, cherrybark oak, hickory, and a fairly dense understory that consisted of American hornbeam, pawpaw, spicebush, and red buckeye.
Community 1.2
Dry-Mesic Mixed Oak – Hickory – Pine Dense Woodland/ForestMany of the same canopy components of Phase 1 occurs in this phase with the possible exception of an increased presence of shade tolerant species entering the community. A major distinction is the crowding and encroachment of an expanding understory into higher strata of the woodland profile. Overall, this community phase supports a greater density or cover of woody vegetation at most all height strata or classes. The herbaceous ground cover is generally the most affected level of the community due to higher shade and loss of growing space.
Pathway 1.1A
Community 1.1 to 1.2This pathway represents a decrease in disturbance frequency leading to a more closed, late development system. Overall, disturbance is light, infrequent, and localized – the result of single tree senescence or small group windthrow and a suppressed fire return interval.
Pathway 1.2A
Community 1.2 to 1.1This pathway involves an increase in disturbance. Mixed severity fire is anticipated to thin this community back to Phase 1.1. Additional disturbances include larger gap- to incomplete stand-scale openings due to wind, ice, and forest management (e.g., group selection, basal area reductions).
State 2
Post Large-scale Disturbance Forest StateThis state is characterized by the regeneration or regrowth of a pre-existing forest stand following a major, stand-replacing disturbance. Scale of the disturbance is at the stand level and is greater than one acre (Johnson et al., 2009). Potential disturbances include catastrophic windstorms, severe fire, insect outbreaks, silvicultural clearcuts, and particularly destructive ice storms. The resulting, even-aged stand (or single-cohort) is set on a new course of development, which is highly dependent upon several critical factors including: the composition and structure of the stand prior to the disturbance; the degree or intensity of the disturbance; size and configuration of the disturbed area; and distance to seed sources.
Composition and condition of the stand prior to a major disturbance may dictate, in large part, future composition of the regenerating stand. Although colonization by new species is expected soon after the disturbance, many of the pre-existing overstory components are anticipated to occupy position in the new, developing stand – their presence arising mainly from stump or root sprouts, advance regeneration, and germination from the seed bank (Oliver and Larson, 1990).
If the intensity of the disturbance only removed the overstory and damage to the understory strata was light, then understory components of advance regeneration may proliferate in the new opening. This may be a desired condition if managing for an oak shelterwood harvest and subsequent oak recruitment. However, this scenario is particularly problematic in high-graded stands.
Community 2.1
Woodland/Forest RegrowthSoon after overstory removal, numerous species may colonize large openings and influence the dynamics of the site. Initial colonizers are often forbs, graminoids, and vines that may have existed in the seed bank, were forest floor components prior to disturbance, or transported into the site via wind and/or animals. Early successional or pioneer species may include winged elm, sumac, greenbrier, grapevine, blackberry, and various graminoids. Overstory species anticipated to occur during the stand-initiation stage include post oak, southern red oak, black oak, shortleaf pine, and various hickories on the driest sites and white oak, northern red oak, hickory, beech, maple on moist, protected sites. Composition of the young stand will vary dramatically if the disturbance is a well-designed and implemented shelterwood harvest that favors the advancement of an established oak understory.
For stands that were highly altered prior to the disturbance (e.g., high-graded), intensive management may be necessary in order to establish a desired composition. Management actions may include controlling undesirable species mechanically and chemically and planting the desired components.
State 3
High-graded/Grazed ForestForests in this state have undergone repeated select harvests over time. Actions leading to this condition consist of removing the biggest and best trees of the most desirable species and leaving low-quality trees (damaged and deformed) and undesirable species. This action, conducted repeatedly, can cause tremendous shifts in species composition and can decrease the vigor and health of the residual stand. Without implementing carefully prescribed management actions, species composition of extreme high-graded stands may remain in a highly altered condition for many decades, even after large, stand-replacing disturbances resets “successional opportunity.”
Today, this vegetation state probably represents the conditions of many forest stands throughout the distribution of this site. Local stands in which desirable species such as oaks and shortleaf pine were repeatedly targeted often results in sites with proportionally more hickory.
Because “overgrazed woods” often consists of components very similar to high-graded stands, uncontrolled livestock access to forests is also included in this state. This does not take into account carefully prescribed and/or managed forms of forest grazing (e.g., agroforestry or silvopasture), which generally has a mutual goal of providing quality forage and productive forest management. The conditions considered and represented here are the extreme cases of long-term forest grazing; this form of uncontrolled access has been referred to as “turning livestock into the woods” (Brantly, 2014).
A single community phase is selected to represent the breadth of conditions that may be anticipated in stands having been high-graded and uncontrolled access by livestock.
Community 3.1
Hickory-Oak/HophornbeamHigh-graded stands generally consist of a paucity of oaks. Species typically left or avoided during harvests often include hickory and practically the entire understory. This has resulted in canopies largely comprised of the preceding species along with a dense understory of hophornbeam and “scrub oak” or undesirable species such as post oak and blackjack oak. Noticeable characteristics of this condition are a conspicuous reduction of more merchantable oaks and other valuable hardwoods.
The most palatable forage of a forest stand is typically the herbaceous understory, which is targeted first. The combined effects of trampling, browsing woody plants, and foraging on the herbaceous layer often results in a high percentage of bare soil, exposed roots, and an open understory. Furthermore, overstory trees occurring in stands with high livestock traffic grow more slowly over time (Johnson, 1952).State 4
Timber ManagementThis state represents the breadth of forest management activities on this site. Various management or silvicultural methods can lead to very different structural and compositional results within a managed stand. The range of methods are diverse and include even-aged (e.g., clearcut and shelterwood) and uneven-aged (single tree, diameter-limit, basal area, group selection, etc.) approaches. Included within these approaches is an option to use disturbance mechanisms (e.g., fire, TSI, etc.) to reduce competition and achieve maximum growth potential of the desired species. Inherently, these various approaches result in different community or “management phases” and possibly alternate states. The decision to represent these varying approaches and management results into a single state and phase at this time hinges on the need for additional information in order to formulate definitive pathways, management actions, and community responses. Forthcoming inventories and description iterations of this site will provide more detail on this state and associated management phases.
Community 4.1
Mixed Oak – Pine (Various)Some of the most desirable timber consists of oak and tall mixed hardwoods on protected sites and shortleaf pine on the drier sites. Depending on the desired end product, management activities will differ. Management for oak dominant stands may be achieved by shelterwood and/or seed tree approaches. Managing for shortleaf pine may only require timber stand improvement methods where pine is currently dominant, or artificial regeneration may be called for where other hardwoods predominate. The droughty portions of this site respond well to fire, and low intensity ground fires on a frequent return interval can be an effective tool for reducing competition and potentially enhancing production of individual trees. Conversely, competition intensifies from various hardwoods on more moist sites and managing for oak can be problematic (see Johnson et al., 2009).
The complex distribution of soils on this site may affect the response of a given stand. For this very reason, consideration of site factors and conditions should be applied into the decision-making process well before management begins. Finding the appropriate approach for a given stand and environment necessitates close consultation with trained, experienced, and knowledgeable forestry professionals. It is strongly urged and advised that professional guidance be secured and a well-designed silvicultural plan developed in advance of any work conducted.State 5
Grassland/PasturelandThis state is representative of sites that have been converted to and maintained in pasture and forage cropland, typically a grass – legume mixture. For pastureland, planning or prescribing the intensity, frequency, timing, and duration of grazing can help maintain desirable forage mixtures at sufficient density and vigor (USDA-NRCS, 2010; Green et al., 2006). Overgrazed pastures can lead to soil compaction and numerous bare spots, which may then become focal points of accelerated erosion and colonization sites of undesirable plants or weeds.
Establishing an effective pasture management program can help minimize the rate of weed establishment and assist in maintaining vigorous growth of desired forage. An effective pasture management program includes: selecting well-adapted grass and/or legume species that will grow and establish rapidly; maintaining proper soil pH and fertility levels; using controlled grazing practices; mowing at proper timing and stage of maturity; allowing new seedings to become well established before use; and renovating pastures when needed (Rhodes et al., 2005; Green et al., 2006). It is strongly advised that consultation with State Grazing Land Specialists and District Conservationists at local NRCS Service Centers be sought when assistance is needed in developing management recommendations or prescribed grazing practices.
Community 5.1
Select Forage/Species MixtureThis community phase represents commonly planted forage species on pasturelands, haylands, and open grasslands. The suite of plants established on any given site may vary considerably depending upon purpose, management goals, usage, and soils. Most systems include a mixture of grasses and legumes that provide forage throughout the growing season. Cool season forage may include tall fescue (Schedonorus arundinaceus), orchardgrass (Dactylis glomerata), white clover (Trifolium repens), and red clover (T. pratense), and warm season forage often consists of bermudagrass (Cynodon dactylon), bahiagrass (Paspalum notatum), and annual lespedeza (Kummerowia spp.). Several additional plants and/or species combinations may be desired depending on the objectives and management approaches and especially, local soils.
Should active management (and grazing) of the pastureland be halted, this phase will transition to “old field” conditions, which is the transitional period between a predominantly open, herbaceous field and the brushy stage of a newly initiated stand of trees.
Transition T1A
State 1 to 2This pathway represents a large-scale, stand replacing disturbance, which may be caused by a catastrophic windstorm (e.g., straight-line winds, tornado), ice storm, severe fire, or a silvicultural clearcut. For this stressor to occur, most or all of the overstory must be removed or destroyed. A few residual trees may persist, but overall, the disturbance must be intensive enough, at least one acre and larger (Johnson et al., 2009), that a new, even-aged stand is created.
Transition T1B
State 1 to 3Repeated selective harvesting or high-grading over time can cause shifts in species composition, structure, and overall health of affected stands. High-grading occurs when the most desirable trees of select species are repeatedly removed leaving behind inferior, low quality stems and undesirable species. This transition also includes uncontrolled access by livestock and impacts from sustained, selective grazing and browsing.
Transition T1C
State 1 to 4This pathway consists of prescribed silvicultural activities specifically designed to meet stand compositional and production objectives.
Transition T1D
State 1 to 5Actions required to convert forests to pasture or forage production include forest clearing, stump removal, herbicide application, seedbed preparation, and the establishment of desired plants.
Restoration pathway R2A
State 2 to 1This pathway represents a return to reference conditions through natural succession, if the disturbance occurred within a reference community.
Transition T2A
State 2 to 4This pathway represents the development of an even-aged stand that is prescribed to meet compositional and production objectives.
Transition T2B
State 2 to 5Pathway represents a conversion of the emerging stand to pastureland or hayland. Actions required include forest clearing, stump removal, herbicide application, seedbed preparation, and the establishment of desired plants.
Restoration pathway R3A
State 3 to 2This pathway represents a large-scale, stand replacing disturbance, which may be caused by a catastrophic windstorm (e.g., straight-line winds, tornado), ice storm, severe fire, landslide, or a silvicultural clearcut.
Transition T3A
State 3 to 5Actions include forest clearing, stump removal, herbicide application, seedbed preparation, and the establishment of desired plants.
Restoration pathway R4A
State 4 to 1Natural succession over a period of time coupled with disturbance such as low intensity (and possibly mixed severity fire) may transition a former timber-managed stand to one supporting reference conditions. Some question remains whether a return to reference conditions will occur in every situation, especially since some components may have been selectively culled from the stand. Management activities to aide recovery may include exotic species control and silvicultural treatment.
Restoration pathway R4B
State 4 to 2This pathway represents a large-scale, stand-initiating disturbance, which effectively removes most or all of the pre-existing overstory. Disturbances may include a catastrophic windstorm, severe wildfire, and silvicultural management (even-aged).
Transition T4A
State 4 to 3Repeated selective harvesting or high-grading of stands over time can cause shifts in species composition, structure, and overall health of affected stands. This transition also includes uncontrolled access by livestock and impacts from sustained, selective grazing and browsing. Impacts from continual grazing and uncontrolled access can result in the removal of palatable understory components, alteration of species composition in current and future stands, conditions for exotic plant invasions, and soil compaction and erosion.
Transition T4B
State 4 to 5Actions include forest clearing, stump removal, herbicide application, seedbed preparation, and the establishment of desired plants.
Restoration pathway R5A
State 5 to 1This pathway represents natural succession back to perceived reference conditions. The period required for this transition to take place likely varies by location and is dependent upon local site conditions. LANDFIRE models (2008) suggest that over 60 years is required for a return to a late development community and this pathway is highly dependent upon species present in the developing stand in addition to the appropriate level and type of disturbance (e.g., fire return interval). Significant efforts may be required before a return to reference conditions is achieved (e.g., exotic species control, appropriate intensity and return interval of fire, potential artificial regeneration of community components, etc.).
Restoration pathway R5B
State 5 to 4This pathway represents prescribed management strategies for transitioning abandoned pastureland to managed woodland. Activities may include artificial regeneration of desired species; exotic species control; appropriate intensity and return interval of fire.
Additional community tables
Table 5. Community 1.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 6. Community 1.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 7. Community 2.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 8. Community 3.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 9. Community 4.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 10. Community 5.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Interpretations
Supporting information
Other references
Brantly, S. 2014. Forest grazing, silvopasture, and turning livestock into the woods. USDA National Agroforestry Center, Agroforestry Note – 46. 4 p. [Online] Available: http://nac.unl.edu/documents/agroforestrynotes/an46si09.pdf.
Braun, E.L. 1950. Deciduous Forests of Eastern North America. Hafner Press, New York. 596 p.
Call, R.E. 1891. Annual Report of the Geological Survey of Arkansas for 1889. Vol. II. The Geology of Crowley’s Ridge. Woodruff Printing Co., Little Rock, AR. 283 p.
Chapman, S.S., J.M. Omernik, G.E. Griffith, W.A. Schroeder, T.A. Nigh, and T.F. Wilton. 2002. Ecoregions of Iowa and Missouri (color poster with map, descriptive text, summary tables, and photographs): Reston, Virginia, U.S. Geological Survey (map scale 1:1,800,000).
Clark G.T., J.A. Akers, S.W. Bailey, W.H. Freeman, M.H. Hill, L.P. Lowman, S.O. Loyd, W.R. Randel, R.B. Rosen, and J.S. Workman. 1974. Preliminary ecological study of Crowley’s Ridge. In: Arkansas Department of Planning. Arkansas Natural Area Plan. Little Rock, AR. 248 p.
Green, Jonathan D., W.W. Witt, and J.R. Martin. 2006. Weed management in grass pastures, hayfields, and other farmstead sites. University of Kentucky Cooperative Extension Service, Publication AGR-172.
Johnson, E.A. 1952. Effect of farm woodland grazing on watershed values in the southern Appalachian Mountains. Journal of Forestry 50 (2): 109-113.
Johnson, P.S., S.R. Shifley, and R. Rogers. 2009. The Ecology and Silviculture of Oaks. 2nd Edition. CABI, Cambridge, MA. 580 p.
LANDFIRE. 2008. LANDFIRE Biophysical Setting Models. Biophysical Setting 45. (2008, February - last update). Homepage of the LANDFIRE Project, U.S. Department of Agriculture, Forest Service; U.S. Department of Interior, [Online]. Available:
http://www.landfire.gov/index.php (Accessed: 1 July 2014).
McNab, W.H.; Cleland, D.T.; Freeouf, J.A.; Keys, Jr., J.E.; Nowacki, G.J.; Carpenter, C.A., comps. 2005. Description of ecological subregions: sections of the conterminous United States [CD-ROM]. Washington, DC: U.S. Department of Agriculture, Forest Service. 80 p.
NatureServe. 2011. NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. NatureServe, Arlington, Virginia. Available: http://www.natureserve.org/explorer. (Accessed: February 9, 2011).
Nelson, P. 2005. The Terrestrial Natural Communities of Missouri. Third edition. Missouri Natural Areas Committee, Department of Natural Resources and the Department of Conservation, Jefferson City, MO. 550 p.
Oliver, C.D. and B.C. Larson. 1990. Forest Stand Dynamics. McGraw Hill, Inc., New York, NY. 476 p.
Rhodes, G.N., Jr., G.K. Breeden, G. Bates, and S. McElroy. 2005. Hay crop and pasture weed management. University of Tennessee, UT Extension, Publication PB 1521-10M-6/05 (Rev). Available: https://extension.tennessee.edu/washington/Documents/hay_crop.pdf.
[USDA-NRCS] United States Department of Agriculture, Natural Resources Conservation Service. 2006. Land Resource Regions and Major Land Resource Areas of the United States, the Caribbean, and the Pacific Basin. U.S. Department of Agriculture Handbook 296.
[USDA-NRCS] United States Department of Agriculture, Natural Resources Conservation Service. 2010. Conservation Practice Standard: Prescribed Grazing. Practice Code 528. Updated: September 2010. Field Office Technical Guide, Notice 619, Section IV. [Online] Available: efotg.sc.egov.usda.gov/references/public/ne/ne528.pdf.
[USDA-NRCS] United States Department of Agriculture, Natural Resources Conservation Service. 2016. Official Soil Series Descriptions. Available online: https://soilseries.sc.egov.usda.gov/osdname.asp. (Accessed: 17 May 2016).
[USDA-SCS] United States Department of Agriculture, Soil Conservation Service. 1992. Hardwood forest grazing. Woodland Fact Sheet No. 7. Columbia, Missouri. 2 p. [Online] Available: www.forestandwoodland.org/uploads/1/2/8/8/12885556/hardwood_forest_grazing1.pdf.
Woods, A.J., T.L. Foti, S.S. Chapman, J.M. Omernik, J.A. Wise, E.O. Murray, W.L. Prior, J.B. Pagan, Jr., J.A. Comstock, and M. Radford. 2004. Ecoregions of Arkansas (color poster with map, descriptive text, summary tables, and photographs): Reston, Virginia, U.S. Geological Survey (map scale 1:1,000,000).
Contributors
Barry Hart
Approval
Matthew Duvall, 3/24/2025
Acknowledgments
Tom Foti (Ecologist, Arkansas Natural Heritage Commission) and Henry Langston (Arkansas Highway Department) provided invaluable discussion, knowledge, and experience with regards to the soils and vegetation associated with this site.
Rangeland health reference sheet
Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.
Author(s)/participant(s) Contact for lead author Date 08/06/2026 Approved by Approval date Composition (Indicators 10 and 12) based on Annual Production Indicators
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Number and extent of rills:
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Presence of water flow patterns:
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Number and height of erosional pedestals or terracettes:
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Bare ground from Ecological Site Description or other studies (rock, litter, lichen, moss, plant canopy are not bare ground):
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Number of gullies and erosion associated with gullies:
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Extent of wind scoured, blowouts and/or depositional areas:
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Amount of litter movement (describe size and distance expected to travel):
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Soil surface (top few mm) resistance to erosion (stability values are averages - most sites will show a range of values):
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Soil surface structure and SOM content (include type of structure and A-horizon color and thickness):
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Effect of community phase composition (relative proportion of different functional groups) and spatial distribution on infiltration and runoff:
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Presence and thickness of compaction layer (usually none; describe soil profile features which may be mistaken for compaction on this site):
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Functional/Structural Groups (list in order of descending dominance by above-ground annual-production or live foliar cover using symbols: >>, >, = to indicate much greater than, greater than, and equal to):
Dominant:
Sub-dominant:
Other:
Additional:
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Amount of plant mortality and decadence (include which functional groups are expected to show mortality or decadence):
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Average percent litter cover (%) and depth ( in):
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Expected annual annual-production (this is TOTAL above-ground annual-production, not just forage annual-production):
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Potential invasive (including noxious) species (native and non-native). List species which BOTH characterize degraded states and have the potential to become a dominant or co-dominant species on the ecological site if their future establishment and growth is not actively controlled by management interventions. Species that become dominant for only one to several years (e.g., short-term response to drought or wildfire) are not invasive plants. Note that unlike other indicators, we are describing what is NOT expected in the reference state for the ecological site:
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Perennial plant reproductive capability:
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PrintThe Ecosystem Dynamics Interpretive Tool is an information system framework developed by the USDA-ARS Jornada Experimental Range, USDA Natural Resources Conservation Service, and New Mexico State University.
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