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Conservation Service
Ecological site R044AB020MT
Gravelly (Gr)
LRU 44A-B
Last updated: 3/08/2022
Accessed: 05/30/2026
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Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.
MLRA notes
Major Land Resource Area (MLRA): 044A–Northern Rocky Mountain Valleys
For further information regarding MLRAs, refer to:
http://soils.usda.gov/survey/geography/mlra/index.html
Land Resource Unit (LRU) 44A-B:
• Moisture Phase: xeric - ustic
• Temperature Phase: frigid, cool – frigid, warm
• Dominant Cover: rangeland
• Representative Value (RV) Effective Precipitation: 14-19 inches
• RV Frost Free Days: 70-120 days
Site Concept:
Site does not receive any additional water.
Soils are:
• not saline or saline-sodic.
• not coarse-granular clay.
• moderately deep, deep, or very deep with less than 15% stone and boulder cover.
• sandy-skeletal within 10” of soil surface.Associated sites
R044AB134MT Shallow To Gravel (Swgr) LRU 44A-B
Similar sites
R044AB036MT Droughty (Dr) LRU 44A-B
Loamy skeletal material in 10-20
R044AB134MT Shallow To Gravel (Swgr) LRU 44A-B
Sandy skeletal material in 10-20
Table 1. Dominant plant species
Tree Not specified
Shrub Not specified
Herbaceous (1) Festuca campestris
(2) Festuca idahoensisPhysiographic features
The Gravelly (Gr) ecological site (R044AB020MT) is located within LRU “B” in MLRA “44A.” This ecological site usually occurs on outwash plains, alluvial fans, stream terraces, or moraines. The slope ranges from 0 to 55%. This site occurs on all exposures; effects of aspect can be significant in LRU assignment.
Table 2. Representative physiographic features
Landforms (1) Outwash plain
(2) Stream terrace
(3) Alluvial fan
Elevation 1800 – 5000 ft Slope 0 – 55 % Water table depth 42 – 0 in Aspect E, S, W Climatic features
The dissected Northern Rocky Mountain Valleys of MLRA 44A are considered to have a maritime climate. Precipitation is evenly distributed throughout the year with less than about 35% of the annual precipitation occurring during the growing season in Montana. Rainfall occurs as high-intensity, convective thunderstorms in the spring and fall. Most of the precipitation in the winter is snow or rain on fully or partially frozen ground. Average precipitation for LRU-B is 17”, and the frost-free period averages 95 days.
See Climatic Data Sheet for more details (Section II of the Field Office Technical Guide: http://efotg.nrcs.usda.gov/efotg_locator.aspx?map=MT) or reference the following climatic Web site: http://www.wrcc.dri.edu/climsum.html.Table 3 Representative climatic features
Frost-free period (average) 100 days Freeze-free period (average) 120 days Precipitation total (average) 20 in BarLineFigure 1. Monthly precipitation range
BarLineFigure 2. Monthly average minimum and maximum temperature
Figure 3. Annual precipitation pattern
Figure 4 Annual average temperature pattern
">Influencing water features
Soil features
These soils are typically very deep, well-drained soils that formed in alluvium and outwash deposits. Soil consists of a sandy-skeletal soil material (averages > 35% rock fragments by volume within 10” of surface). This skeletal material and sandy textures significantly decrease the water- holding capacity of the ecological site. Skeletal soil material may or may not be present to the surface. Surface textures (< 2 mm) usually range from loamy sand to loam and are typically gravelly or very gravelly.
Table 4. Representative soil features
Surface texture (1) Loam
(2) Sandy loam
(3) Fine sandy loam
Family particle size (1) Sandy
Drainage class Well drained Permeability class Moderate to moderately rapid Soil depth 20 – 0 in Available water capacity
(0-40in)1 – 2.5 in Calcium carbonate equivalent
(0-40in)0 – 20 % Electrical conductivity
(0-40in)0 – 1 mmhos/cm Sodium adsorption ratio
(0-40in)0 – 12 Soil reaction (1:1 water)
(0-40in)6.3 – 8.2 Ecological dynamics
The Gravelly ecological is characterized by the production and composition of plant species in the Reference Plant Community, which is defined by soils, precipitation, and the temperature regime influencing the site. The presumed Reference Plant Community type of this site is dominated by cool-season perennial bunchgrass species, primarily rough fescue (Festuca campestris) with minor components of perennial forbs and low-growing shrubs. LRU-B occurs in the valleys of western Montana, on rangelands with a xeric and ustic soil moisture phase, a frigid, cool to frigid, warm soil temperature phase, 15-19” of effective precipitation, and between 70 and 120 consecutive frost-free days annually. This site is characterized by moderately deep to very deep, coarse-textured to medium-textured surface horizons, and are sandy-skeletal to within 10” of the soil surface.
The majority of precipitation comes early in the form of snow and spring rain. Summers are usually dry. The growing season is short and cool; primary growth typically occurs between May and July, and dominant plants are those that have adapted to these conditions.
In response to disastrous fires in 1910, new firefighting policies were established. Wildland fire suppression became an important driving factor in the ecology of western rangelands. Livestock grazing during the late 1800s and early 1900s often occurred at very heavy levels. Heavy grazing resulted in a severe reduction in fine fuels, which further reduced potential for natural fires. These two actions altered the natural fire interval.
Fire suppression, along with fine-fuels reduction, has interfered with the natural fire interval; many areas have not burned for over 100 years (Arno and Gruell 1986). Prior to 1900, the average natural fire return intervals were probably shorter than 35 years for this MLRA. Historic fire frequency may have ranged from 15 to 75 years. Trees and non-sprouting shrubs were restricted to small patches or widely spaced plants. Following fire on medium-textured soils, perennial bunchgrasses apparently recovered in a few years and were present to fuel subsequent fires, which suppressed woody species and kept them as a minor component of the community (Arno and Gruell 1983).
Historical records indicate, prior to the introduction of livestock (cattle and sheep) during the late 1800s, elk and bison grazed of this ecological site. Evidence shows periodic use by bison was in large numbers and concentrations (Lesica and Cooper 1997). Forage for livestock was noted as minimal in areas recently grazed by bison (Lesica and Cooper 1997).
Significant livestock grazing has occurred on most of this ecological site in western Montana for more than 100 years (beginning with the 1860s gold boom and subsequent settlement through 1900). Indian horse herds were present and numerous for several hundred years prior. The primary type of European livestock grazed in this region has historically transitioned between sheep and cattle with early grazing (pre-1890) dominated by the cattle industry. In the 1890s Montana sheep production began to increase dramatically (> 400%) and dominated the cattle industry for approximately four decades. By the 1930s, livestock production once again favored the cattle industry, which continues to dominate livestock grazing in the region today (Wyckoff and Hansen 2001). The Gravelly ecological site is relatively accessible and many examples were subject to heavy and/or season-long grazing until 1970 or later.
Invasive species are an important part of the ecology of MLRA 44A. Notable invasive species include spotted knapweed (Centaurea stoebe), leafy spurge (Euphorbia esula), sulphur cinquefoil (Potentilla recta), and cheatgrass (Bromus tectorum). Most sites in MLRA 44A are impacted by these invasives. Sites either are currently invaded or have been treated to kill invasives, which reduces the production and changes the composition of forbs and shrubs. Even where invasives are not present, the threat of invasion drives management of this site.
Although there is considerable qualitative experience supporting the pathways and transitions within the State and Transition Model (STM), no quantitative information exists that specifically identify threshold parameters between grassland types and invaded types in this ecological site. For information on STMs, see the following citations: Bestelmeyer el al. 2003, Bestelmeyer et al. 2004, Bestelmeyer and Brown 2005, Stringham et al. 2003.
Rangeland Health Reference Worksheets have been posted for this site on the Montana NRCS Web site (www.mt.nrcs.usda.gov) in Section II of the eFOTG under (F) Ecological Site Descriptions (ESD).
Plant Communities and Transitional Pathways
A State and Transition Model for the Gravelly ecological site (44AB020MT) is depicted in Figure 1. Thorough descriptions of each state, transition, plant community, and pathway follow the model. This model is based on available experimental research, field observations, and interpretations by experts and is likely to change as knowledge increases.
Plant communities will differ across the MLRA because of the naturally occurring variability in weather, soils, and aspect. The biological processes on this site are complex; therefore, representative values are presented in a land management context. The species lists are representative and are not botanical descriptions of all species occurring, or potentially occurring, on this site. The species lists are not intended to cover every situation or the full range of conditions, species, and responses for the site.
Both percent species composition by weight and percent canopy cover are used in this ESD. Most observers find it easier to visualize or estimate percent canopy for woody species (trees and shrubs). Canopy cover drives the transitions between communities and states because of the influence of shade and interception of rainfall. Species composition by dry weight remains an important descriptor of the herbaceous community and of the community as a whole. Woody species are included in species composition for the site. Calculating similarity index requires use of species composition by dry weight.
State and transition model
Custom diagramStandard diagram
Figure 5. 44AB020MT Gravelly
More interactive model formats are also available. View Interactive Models
More interactive model formats are also available. View Interactive Models
Click on state and transition labels to scroll to the respective textState 1 submodel, plant communities
State 2 submodel, plant communities
State 3 submodel, plant communities
State 1
Taller Bunchgrass StateThis state is characterized by cool-season bunchgrasses and is represented by two communities that differ mainly in the percent composition rough fescue and Idaho fescue (Festuca idahoensis). Shrubs and forbs are a minor component in this state.
Community 1.1
Reference Plant Community - Taller Bunchgrass Community
Figure 6. 44AB Gravelly 1
Figure 7. 44AB Gravelly 2
Figure 8. 44AB Gravelly 3
The Taller Bunchgrass Community (1.1) is dominated by rough fescue, a taller cool-season bunchgrass with a minor component of forbs and low-growing shrubs. Rough fescue is typically the dominant producer in the Taller Bunchgrass Community (1.1), while Idaho fescue and bluebunch wheatgrass (Pseudoroegneria spicata) are subdominant. Many common forb species exist on this site, including silky lupine (Lupinus sericeus). Shrub species, including Woods’ rose (Rosa woodsii), are present as a minor part of the community.
The Taller Bunchgrass State generally occurs on the Gravelly site in areas where proper grazing management practices have been implemented over a long period. The Taller Bunchgrass Community can be maintained through the implementation of properly managed grazing that provides adequate growing-season deferment to allow establishment of taller grass propagules and/or the recovery of vigor of stressed plants.
Taller Bunchgrass Community (1.1) in general is resistant to change with proper grazing management and near normal precipitation. However, rough fescue and bluebunch wheatgrass lack resistance to grazing during the spring growing season. Subdominant species, such as Idaho fescue and needleandthread, tolerate higher grazing pressure, and may increase in cover under prolonged drought conditions. This increase drives the community shift to the Mixed Bunchgrass Plant Community (1.2).
The Taller Bunchgrass Community is moderately resilient. This community will return to dynamic equilibrium (1.2A) following a relatively short period of stress, such as drought or short-term overgrazing, provided the return of favorable or normal growing conditions occurs along with implementation of proper grazing management. This equilibrium will occur if canopy cover did not fall below 50%, and rough fescue did not fall below 10% of species composition.
Rough fescue and bluebunch wheatgrass lack resistance to grazing during the critical growing period: spring. They may decline in vigor and production if grazed in the spring more than one year in three (Mengli et al. 2005, McLean and Wikeem 1985, Wilson et al. 1960).
Periodic fire increases the resilience of the Taller Bunchgrass Community (1.1) by reducing competition and canopy cover of less fire-tolerant species. Fire also removes decadent herbaceous material, particularly from taller bunchgrasses, which promotes increased vigor and seedling establishment. Timing and intensity of a fire are critical components that can have varying positive or negative effects on this plant community. Fire does increase risk of invasion from invasive species, most notably cheatgrass. At least two growing seasons of rest are recommended to allow plants to recover after fire.
Increaser species on this site are generally endemic species released by disturbance. These subdominant species of grasses, forbs, and shrubs are more tolerant to grazing pressure than rough fescue and bluebunch wheatgrass. Improper grazing management can reduce plant vigor of rough fescue, which can lead to reduced plant size or plant death. Species with higher grazing tolerance will increase in production as they use resources made available by the decrease in rough fescue. Improper grazing management can also lead to degraded soil properties through compaction, erosion, decrease in organic matter, and increase in exposure because of reduction in litter cover. Idaho fescue is not only more tolerant to higher grazing pressure but can also grow on less fertile soils than rough fescue (USDA/NRCS 2007).
Under improper grazing management, the Reference Plant Community shifts to the Mixed Bunchgrass Community (1.2). If overgrazing continues, invasive grass and forb species can move into the plant community and the site can transition to the Invaded State (3).
While the Taller Bunchgrass Community (1.1) is resilient to degradation under proper management, the community remains at risk of invasion by aggressive nonnative species because of the ability of spotted knapweed, leafy spurge, and cheatgrass to invade healthy rangelands and the widespread presence of propagules. Healthy plant communities are most resilient to invasives although many examples exist of well-managed areas that have been invaded by spotted knapweed. Due to the ability of spotted knapweed and other aggressive species to invade any community, all communities, including the Reference Plant Community (1.1) are “at risk communities” to cross the threshold to the Invaded State (3).
Invasives impact plant communities even if the site does not yet have critical populations of invasives. Almost all reference sites had at least trace amounts of spotted knapweed and/or cheatgrass. It is believed that most sites with only trace amounts have been chemically treated for invasives at some point. These treatments would have affected other broad-leafed species (forbs and shrubs). It is likely that this site had more potential for forb and shrub production than found on current reference sites.
Rock cover on the soil surface is minimal and does not impact productivity of this site. Plant basal cover is expected to be about 15 to 25% and bare ground is expected to be < 10%. The soils of this site have high soil stability values.
The following production figures do not represent the lowest or highest possible production for the Reference Plant Community (1.1). For example, the high figure is not the most production that can occur in a wet year in the most mesic portion of the LRU. These values represent the range of variability for each species across the extent of the ecological site. Usually, values in the low production column represent production at the dry end of the LRU and those in the high production column represent production at the wet end of the LRU.
Even the most stable communities exist within a range of dynamic equilibrium of species composition. The following table shows an example of species composition; the example is not the only mix of species possible in the Taller Bunchgrass Community (1.1).
Figure 9. Annual production by plant type (representative values) or group (midpoint values)
Table 5. Annual production by plant type
Plant type Low
(lb/acre)Representative value
(lb/acre)High
(lb/acre)Grass/Grasslike 850 1148 1445 Forb 100 135 170 Shrub/Vine 50 68 85 Total 1000 1351 1700 Figure 10. Plant community growth curve (percent production by month). MT0441 , 44A - Uplands. Includes all upland sites in 44A.
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec J F M A M J J A S O N D 10 25 35 15 10 5 Community 1.2
Mixed Bunchgrass CommunityIdaho fescue tolerates grazing pressure better than rough fescue. It increases in species composition when more palatable and less grazing tolerant plants decrease due to improper grazing management. Idaho fescue and rough fescue share dominance in the Mixed Bunchgrass Community (1.2). Bluebunch wheatgrass is subdominant. Other subdominant grass species that are more tolerant to grazing are likely to increase include Sandberg bluegrass (Poa secunda) needleandthread (Hesperostipa comata), prairie junegrass (Koeleria macrantha) and Kentucky bluegrass (Poa pratensis). Some increaser forbs species may include silky lupine (Lupinus sericeus), field chickweed (Cerastium arvense), ballhead sandwort (Arenaria congesta), northern bedstraw (Galium boreale) and pussytoes (Antennaria spp.). Fringed sagewort, Woods’ rose (Rosa woodsii) and common snowberry (Symphoricarpus albus) are shrubs that also increase under prolonged drought or heavy grazing.
Heavy continuous grazing will reduce plant cover, litter, and mulch. Bare ground will increase and expose the soil to erosion. Litter and mulch will move off-site as plant cover declines. As long as the canopy cover remains > 50% and production of rough fescue is > 10% of total biomass production, the site can return to the Taller Bunchgrass Community (Pathway 1.2A) under proper grazing management and favorable growing conditions.
Idaho fescue will continue to increase in dominance until it makes up 80% or more of species composition. Once rough fescue has been reduced on the site to < 10% and canopy cover decreased to below 50%, it may be difficult for the site to recover to the Reference Plant Community (1.1). The risk of soil erosion increases when canopy cover decreases to below 50%. As soil properties degrade there will be loss of organic matter, reduced litter, compaction, and reduced soil fertility. Degraded soil properties increase the difficulty of reestablishing rough fescue plants and returning to the Reference Plant Community (1.1).
The Mixed Bunchgrass Community (1.2) is the “At-Risk” Plant Community for this ecological site. When overgrazing continues, increaser species such as Idaho fescue, needleandthread and native forb species will become more dominate and this triggers the change to the Altered Bunchgrass State (2) or the Invaded State (3). Until the Mixed Bunchgrass Community (1.2) crosses the threshold into the Idaho Fescue Community (2.1) or the Invaded Community (3.1), this community can be managed toward the Rough Fescue Community (1.1) using prescribed grazing and strategic weed control. It may take several years to achieve this recovery, depending on growing conditions, vigor of remnant rough fescue plants, and the aggressiveness of the weed treatments.Pathway 1.1A
Community 1.1 to 1.2Rough fescue loses vigor when overgrazed. When vigor declines enough for plants to die or become smaller, species with higher grazing tolerance (most often Idaho fescue) increase in vigor and production as they use the resources previously used by rough fescue and bluebunch wheatgrass. Decrease of species composition by weight of rough fescue to < 50% indicates that the plant community has shifted to the Mixed Bunchgrass Community (1.2). The driver for this community pathway 1.1A is improper grazing management. This shift is triggered by the loss of vigor of rough fescue.
Pathway 1.2A
Community 1.2 to 1.1The Mixed Bunchgrass Community (1.2) will return to the Taller Bunchgrass Community (1.1) with proper grazing management that provide sufficient critical growing season deferment in combination with proper grazing intensity. Favorable moisture conditions will facilitate or accelerate this transition. The driver for this community shift (1.2A) is the increase in vigor of rough fescue to the point that it represents more than 50% of species composition. The trigger for this shift is the change in grazing management that favors rough fescue.
State 2
Alternate Bunchgrass StateThis state is characterized by having < 10% rough fescue and < 50% canopy cover. State 2 is represented by two communities that differ in the percent composition of Idaho fescue, production, and soil degradation. Production in this state is considerably lower than in the Taller Bunchgrass State (1). Some native plants tend to increase under prolonged drought and/or heavy grazing practices. A few of these species include Idaho fescue, needleandthread, Sandberg bluegrass, silky lupine, field chickweed, ballhead sandwort, common snowberry, Wood’s rose and fringed sagewort.
Community 2.1
Idaho Fescue CommunityLong-term grazing mismanagement with continuous growing-season pressure will reduce total productivity of the site and lead to an increase of bare ground. Once plant cover is reduced, the site is more susceptible to erosion and degradation of soil properties. Soil erosion or reduced soil fertility will create reduced plant production. This soil erosion or loss of soil fertility indicates the transition to the Altered Bunchgrass State (2), because it creates a threshold that requires input of energy to return to the Taller Bunchgrass State (1). The transition to Idaho Fescue Community (2.1) may be exacerbated by extended drought conditions.
Idaho fescue dominates the Idaho Fescue Community (2.1). Rough fescue makes up < 10% of species composition by dry weight and the remaining rough fescue plants tend to be scattered and low in vigor. Increaser and invader species will become more common and will create more competition for rough fescue in the community. This competition makes it difficult for rough fescue to increase with simply a change in grazing management alone. Therefore, an input of energy will be required for the community to return to the Taller Bunchgrass State (1). Proper grazing management over a longer period of time is a successful strategy to increase cover and production of rough fescue and bluebunch wheatgrass. Canopy cover decreases compared to the Mixed Bunchgrass Community (1.2) < 50%. Wind and water erosion may be eroding soil from the plant interspaces. Soil fertility is reduced; soil compaction is increased; and resistance to soil surface erosion has declined compared to the Taller Bunchgrass State.
This community has crossed a threshold compared to the Mixed Bunchgrass Community (1.2) due to the erosion of soil, loss of soil fertility, or degradation of soil properties which causes a critical shift in the ecology of the site. The affects of soil erosion can alter the hydrology, soil chemistry, soil microorganisms, and soil physics to the point where intensive restoration is required to restore the site to another state or community. Simply changing grazing management cannot create sufficient change to restore the site within a reasonable time frame. Restoration will require a considerable input of energy to move the site back to the Taller Bunchgrass State (1). This state has lost soil or vegetation attributes to the point that recovery to the Taller Bunchgrass State (1) will require reclamation efforts, i.e., soil rebuilding, intensive mechanical treatments, and/or reseeding.
The transition to this community could occur because of overgrazing (often because of failure to adjust stocking rates in response to declining forage production because of increased dominance of unpalatable invasive species), long-term lack of fire, warming climate, or extensive drought. If heavy grazing continues, plant cover, litter, and mulch will further decrease and bare ground will further increase exposing the soil to accelerated erosion. Litter and mulch will move off-site as plant cover declines. The Idaho Fescue Community will then shift to a Sparsely Vegetated Community (2.2). Introduction or expansion of invasive species will further drive the plant community to the Invaded State (3).
Community 2.2
Sparsely Vegetated CommunityVery sparse plant cover and soil surface erosion characterize this community. Grass and forb cover may be very sparse or clumped (canopy cover < 25%). Weeds, annual species, or shortgrass species dominate the plant community. Mid-stature perennial bunchgrass species (e.g., Idaho fescue) may exist, but only in patches.
In this community phase there may be a significant amount of bare ground, and large gaps may occur between plants. Potential exists for soils to erode to the point that irreversible damage may occur. If further soil erosion occurs, there will be a critical negative shift in the ecological processes of this site. Soil erosion combined with lack of organic matter deposition because of sparse vegetation creates changes to the hydrology, soil chemistry, soil microorganisms, and soil physics to the point where intensive restoration is required to restore the site to another state or community. Simply changing management (i.e., improving grazing management) cannot create sufficient change to restore the site within a reasonable period.
This plant community may be in a terminal state that will not return to the reference state because of degraded soil properties and loss of higher successional native plant species.Pathway 2.1A
Community 2.1 to 2.2With continued overgrazing, bunchgrasses and perennial forbs can decrease in the Idaho Fescue Community (2.1) site. Loss of larger bunchgrasses and rhizomatous grasses will increase bare soil and allow increased soil erosion. This shift is frequently accompanied by decreased soil fertility and diminished soil properties. Decreased plant vigor drives this shift. This shift is triggered by continued overgrazing or extended drought in an Idaho Fescue Community (2.1) with poor vigor. Lack of mid-stature bunchgrasses and low production indicates a community shift to the Sparsely Vegetated Community (2.2).
Pathway 2.2A
Community 2.2 to 2.1If a Sparsely Vegetated Community (2.2) is properly managed for several years and growing conditions are favorable, annual production of perennial bunchgrasses and rhizomatous grasses may increase over time and the site may shift back to the Idaho Fescue Community (2.1). The driver for this shift is increased vigor of bunchgrasses and rhizomatous grasses. The trigger is improved grazing management and growing conditions over a long period.
State 3
Invaded StateThe single community described below characterizes this state.
Community 3.1
Invaded CommunityThe Invaded State (3) is characterized by > 25% of invasive species with spotted knapweed, leafy spurge, sulphur cinquefoil, and/or cheatgrass are the dominant invasive species in MLRA 44A. Introduced exotic plant species have been identified as one of the greatest threats to the integrity and productivity of native rangeland ecosystems and conservation of indigenous biodiversity (DiTomaso 2000; Mack et al. 2000). In addition to environmental consequences, damages caused and costs incurred to control invasive plants are several billion dollars each year in the United States (Pimentel et al. 2000).
Invasives are the driving factor throughout western Montana and they are a focal part of the ecology of MLRA 44A. Their ability to take over and dominate a site has become a big concern. Improper grazing management has contributed to the spread of these species.
The potential for altered ecosystem structure and function is high in the Invaded State (3) and can occur in many ways. The increase in invasive species, especially noxious weeds, can lead to reduction of the native bunchgrasses and an increase in the proportion of bare ground, which often results in reduced infiltration rates and increased surface runoff and erosion. Invasion by cheatgrass reduces above and below ground biomass (Ogle et al. 2003), increases plant litter, changes plant community canopy architecture (Belnap and Phillips 2001), reduces soil biota richness and abundance, reduces plant community richness (Belnap et al. 2005), increases wildfire frequency (Whisenant 1990), and potentially facilitates invasion by other noxious or invasive plants. Dense populations of invasive species can cause soil loss to increase because of lack of surface cover (Lacey et al. 1989).
Early in the invasion process there is a lag phase where invasive plant populations remain small and localized before expanding exponentially (Hobbs and Humphries 1995). Based on research conducted in noxious weed-invaded plant communities in Montana, it is reasonable to estimate that 25% dry-weight composition of invasive plant species is the point in the invasion process where spread and abundance is increasing exponentially and where a plant community has crossed a threshold (Masters and Sheley 2001). For aggressive invasive species (i.e., spotted knapweed), this threshold could be < 10%.
Once invasive species dominate the site, either in species composition by weight or in their impact on the community, the threshold has been crossed to the Invaded State (3). Once invasive species such as spotted knapweed, cheatgrass, and leafy spurge become established, they are very difficult to eradicate. Therefore, considerable effort should be placed in preventing plant communities from crossing a threshold to the Invaded State (3) through early detection and proper management. Preventing new invasions is by far the most cost-effective control strategy and typically places an emphasis on education. Control measures used on the noxious plant species impacting this ecological site include chemical, biological, and cultural control methods. The best success has been found with an integrated weed management strategy that incorporates one or several of these options along with education and prevention efforts (DiTomaso 2000).
Production in the invaded community may vary greatly. A site dominated by spotted knapweed, where soil fertility and chemistry remain near potential, may have production near that of the reference community. A site with degraded soils and infestation of cheatgrass may produce only 10-20% of the reference community.
Invasive plant species have effective reproductive strategies, long seed viability in the soil seed bank, and/or allelopathic properties (Williamson and Fitter 1996). Spotted knapweed has allelopathic properties whereby its roots exude catechin, which may limit the growth and establishment of other plant species (Callaway and Vivanco 2007; Bais et al. 2002), thus promoting its own success. An in-vitro experiment showed that other weeds like Dalmatian toadflax (Linaria dalmatica), kochia (Kochia scoparia), diffuse knapweed (Centaurea diffusa), and crops, such as wheat (Triticum aestivum), showed mortality on the fourteenth day after addition of root exudates from spotted knapweed (Bais et al. 2002). This allelopathic property creates highly resilient communities.
Cheatgrass has the ability to establish rapidly and attain community dominance following disturbances such as wildfire (Young and Evans 1978) or other disturbances that create bare soil. Cheatgrass is a successful invader because it has the ability to respond rapidly to increases in resource availability (Norton et al. 2004; Lowe et al. 2003) as well as to compete for water (Pellant 1996).
Cheatgrass was introduced into the United States in packing materials, ship ballast, and likely as a contaminant of crop seed. Cheatgrass was first found in the United States near Denver, Colorado, in the late 1800s. In the late 1800s and early 1900s, cheatgrass spread explosively in the ready-made seedbeds prepared by the trampling livestock hooves of overstocked rangelands. Cheatgrass has developed into a severe weed in several agricultural systems throughout North America, particularly western pastureland, rangeland, and winter wheat fields (NRCS 2009). Today, cheatgrass is found in most of the western states having reached its range of current distribution by 1930. In fact, a survey of 11 western states showed that cheatgrass was present on at least 60 million acres (Pellant 1996).
After arriving in 1893 on the San Juan Islands in Washington, spotted knapweed had established in over 24 counties in three northwestern states by 1924, with several large infestations near Missoula, Montana (Sheley et al. 2005). By 1975, spotted knapweed had spread into most of the western counties of Montana, and today it is found in every county in Montana.
Leafy spurge, a native to Eurasia, was sighted in Park County, Montana as early as 1925 and has since been found in every county in Montana. Overgrazing by livestock has contributed to the spread of leafy spurge (Sheley et al. 2005).Transition T1A
State 1 to 3The Taller Bunchgrass State (1) transitions to the Altered Bunchgrass State (2) if plant canopy cover declines to < 50% and rough fescue decreases to below 10% by dry weight. The trigger for this transition is the loss of taller bunchgrasses, which creates open spots of bare soil. Soil erosion is accompanied by decreased soil fertility driving the transitions to the Altered Bunchgrass State. There are several other key factors signaling the approach of transition T1A: increases in soil physical crusting, decreases in cover of cryptogamic crusts, decreases in soil surface aggregate stability and/or evidence of erosion including water flow patterns, development of plant pedestals, and litter movement. The driver for this transition is improper grazing management and/or long-term drought leading to a decrease in rough fescue composition to < 10%.
Restoration pathway R2A
State 2 to 1The Altered Bunchgrass State (2) has lost soil or vegetation attributes to the point that recovery to the Taller Bunchgrass State (1) will require reclamation efforts, such as soil rebuilding, intensive mechanical treatments, and/or revegetation. The drivers for this restoration pathway are reclamation efforts and proper grazing management. The trigger is restoration efforts.
Transition T2A
State 2 to 3Invasive species can occupy the Altered Bunchgrass State (2) and drive it to the Invaded State (3). The Altered Bunchgrass State is at risk of this transition occurring if invasive propagules are present. The driver for this transition is the presence of critical population levels (> 25%) of invasive species. The trigger is the presence of propagules of invasive species.
Restoration pathway R3A
State 3 to 1Restoration of the Invaded State (3) to the Taller Bunchgrass State (1) requires substantial energy input. The drivers for this restoration pathway are removal of invasive species, restoration of native bunchgrass species, ongoing management of invasives, and proper grazing management. Without maintenance, invasive species are likely to return (probably rapidly) because of the presence of propagules in the soil and increases soil disturbance. The drivers for this reclamation pathway are treatments to reduce or remove invasive/noxious species in combination with favorable growing conditions.
Restoration pathway R3B
State 3 to 2If invasive species are removed without sufficient remnant populations of reference community species (particularly rough fescue), a site in the Invaded State (3) is likely to return to the Altered Bunchgrass State (2) instead of the Taller Bunchgrass State (1). The driver for the reclamation pathway is weed management without reseeding. The trigger is invasive species control.
Additional community tables
Table 6. Community 1.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Grass/Grasslike1 Cool Season Bunchgrasses 750–1275 rough fescue FECA4 Festuca campestris 700–1190 – bluebunch wheatgrass PSSP6 Pseudoroegneria spicata 300–510 – Idaho fescue FEID Festuca idahoensis 200–340 – needle and thread HECO26 Hesperostipa comata 100–170 – needlegrass ACHNA Achnatherum 100–170 – 2 Shortgrasses/Rhizomatous Grasses/Grasslikes 100–170 Grass, perennial 2GP Grass, perennial 50–85 – sedge CAREX Carex 50–85 – thickspike wheatgrass ELLA3 Elymus lanceolatus 50–85 – prairie Junegrass KOMA Koeleria macrantha 50–85 – western wheatgrass PASM Pascopyrum smithii 50–85 – Sandberg bluegrass POSE Poa secunda 50–85 – Forb3 Forbs 100–170 Forb, annual 2FA Forb, annual 50–85 – Forb, perennial 2FP Forb, perennial 50–85 – common yarrow ACMI2 Achillea millefolium 50–85 – rosy pussytoes ANRO2 Antennaria rosea 50–85 – pointedtip mariposa lily CAAP Calochortus apiculatus 50–85 – tiny trumpet COLI2 Collomia linearis 50–85 – parsnipflower buckwheat ERHE2 Eriogonum heracleoides 50–85 – blanketflower GAAR Gaillardia aristata 50–85 – hairy false goldenaster HEVI4 Heterotheca villosa 50–85 – western stoneseed LIRU4 Lithospermum ruderale 50–85 – desertparsley LOMAT Lomatium 50–85 – silky lupine LUSE4 Lupinus sericeus 50–85 – thinleaved owl's-clover ORTE2 Orthocarpus tenuifolius 50–85 – beardtongue PENST Penstemon 50–85 – woolly plantain PLPA2 Plantago patagonica 50–85 – slender cinquefoil POGR9 Potentilla gracilis 50–85 – meadow deathcamas ZIVE Zigadenus venenosus 50–85 – Shrub/Vine4 Shrubs 50–85 Shrub (>.5m) 2SHRUB Shrub (>.5m) 25–43 – Woods' rose ROWO Rosa woodsii 25–43 – Table 7. Community 1.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 8. Community 2.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 9. Community 2.2 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 10. Community 3.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Interpretations
Animal community
Livestock grazing is suitable on this site due to the potential to produce high quality forage. This site may be preferred for grazing by livestock, and animals may congregate in these areas, however if slopes are > 15% and distance from water is too great, livestock grazing will be limited. Management objectives should include maintenance or improvement of rangeland health attributes of this ecological site. Careful management of timing, intensity and duration of grazing to minimize grazing re-growth and providing adequate rest is important. Shorter grazing periods and changing season of use during the growing season are recommended for plant maintenance, health and recovery.
Continuous grazing with improper stocking rates throughout the growing season in pastures year after year will be detrimental, will alter the plant composition and production over time, and will result in a transition to the Mixed Bunchgrass Community (1.2) or potentially hasten a change to the Invaded State (3.1). Transition to other states will depend on how well the site is managed over time with grazing animals as well as other circumstances such as weather conditions over a period of time.
The transition to the Mixed Bunchgrass Community (1.2) can be the result of long-term, continuous grazing and/or repeated critical growing season grazing (early season grazing during stem elongation). This transition can also occur due to a combination of overgrazing and drought. Repeated grazing during stem elongation (generally mid-April through mid-June), can have detrimental affects, especially on the taller key bunchgrass species. Repeated spring grazing and/or repeated and prolonged summer grazing depletes stored carbohydrates, resulting in poor vigor of key forage plants over time and eventual death of these cool-season grasses – this can lead to an increase in less desirable native species and/or noxious weeds.
The Mixed Bunchgrass Community (1.2) can occur across the entire ecological site or can occur in a mosaic with higher and/or lower states. This is most notable in areas that attract additional grazing, such as water sources or salting locations.
The Mixed Bunchgrass Community (1.2) is subject to further degradation to the Altered Bunchgrass State (2) or Invaded State (3). Management should focus on grazing management strategies that will prevent further degradation. Forage quantity and/or quality may be substantially reduced compared to the Reference Plant Community.
In the Altered Bunchgrass State, forage production is substantially reduced compared to the Taller Bunchgrass State. Grazing is possible in the Invaded State, but invasive species are generally much less palatable than native grasses and forage production is greatly reduced in this state. Grazing should be carefully managed to avoid soil loss and degradation of soil properties as well as to ensure adequate livestock health.
Prescriptive grazing should be included in a conservation plan to maintain vigor of key native plant species while targeting the invasive species problem. In some instances, carefully targeted grazing (sometimes in combination with other treatments) can reduce or eliminate populations of invasive species.
Distance to drinking water and slope can reduce grazing capacity within a management unit. Adjustments should only be made for the area that is considered necessary for reduction of animal numbers. For example 30% of a management unit may have 25% slopes and distances of > 1 mile from water; therefore the adjustment is only calculated for 30% of the unit (50% reduction on 30% of management unit). The table below is a general guide for ranches in Montana (Ricketts et al. 2004). Fencing, slope length, management, access, terrain and breeds are all factors that can increase or decrease the percent of grazable acres within a management unit. Adjustments should be made that incorporate pasture conditions when calculating stocking rates.Hydrological functions
The water cycle functions best in the Taller Bunchgrass State (1) with good infiltration and deep percolation of rainfall. The water cycle degrades as the vegetation community declines. Rapid rainfall infiltration, high soil organic matter, good soil structure, and good porosity accompany high total ground cover of around 95%. High ground cover reduces raindrop impact on the soil surface, which keeps erosion and sedimentation transport low. Water leaving the site will have minimal sediment load, which allows for high water quality in associated streams. High rates of infiltration will allow water to move below the rooting zone during periods of heavy rainfall. The Taller Bunchgrass Community (1.1) should have no rills or gullies present and drainageways should be vegetated and stable.
Improper grazing management results in a community shift to the Mixed Bunchgrass Community (1.2). This plant community has slightly reduced canopy cover, but bare ground will be < 10%. Therefore, the water cycle is functioning at a level similar to the Taller Bunchgrass Community (1.1). Compared to the Taller Bunchgrass Community (1.1), infiltration rates are slightly reduced and surface runoff is slightly higher.
In the Altered Bunchgrass State (2) and the Invaded State (3), canopy and ground cover are greatly reduced compared to the Taller Bunchgrass State (1), which impairs the water cycle. Infiltration will decrease and runoff will increase because of reduced ground cover, rainfall splash, soil capping, reduced organic matter, and poor structure. Sparse ground cover and decreased infiltration can combine to increase frequency and severity of flooding within a watershed. Soil erosion is accelerated, quality of surface runoff is poor, and sedimentation increases.Recreational uses
This site provides some limited recreational opportunities for hiking, horseback riding, and big game and upland bird hunting. The forbs have flowers that appeal to photographers. This site provides valuable open space.
Wood products
None
Other products
None
Other information
None
Supporting information
Inventory data references
Information presented was derived from NRCS clipping data, literature, field observations (based on one sampled site and observations from numerous others), and personal contacts with range-trained personnel (i.e., professional opinion of agency specialists, observations of land managers, and outside scientists).
Other references
Arno, S. F., and Gruell, G. E. 1983. Fire history at the forest-grassland ecotone in southwestern Montana. Journal of Range Management 36(3): 332-336.
Arno, S. F., and Gruell, G. E. 1986. Douglas-fir encroachment into mountain grasslands in southwestern Montana. Journal of Range Management 39(3): 272-275.
Bais, H. P., T. S. Walker, F. R. Stermitz, R. H. Hufbauer, and J. M. Vivanco. 2002. Enantiomeric-dependent phytotoxic and antimicrobial activity of (±)-catechin. A rhizosecreted racemic mixture from spotted knapweed. Plant Physiology 128: 1173-1179.
Belnap, J., and S. L. Phillips. 2001. Soil biota in an ungrazed grassland: response to annual grass (Bromus tectorum) invasion. Ecological Applications 11:1261-1275.
Belnap, J., S. L. Phillips, S. K. Sherrod, and A. Moldenke. 2005. Soil biota can change after exotic plant invasion: does this affect ecosystem processes? Ecology 86:3007-3017.
Bestelmeyer, B., and J. R. Brown. 2005. State-and-transition models 101: a fresh look at vegetation change. The Quivira Coalition Newsletter, Vol. 7, No. 3.
Bestelmeyer, B., J. R. Brown, K. M. Havstad, B. Alexander, G. Chavez, J. E. Herrick. 2003. Development and use of state and transition models for rangelands. Journal of Range Management 56(2):114-126.
Bestelmeyer, B., J. E. Herrick, J. R. Brown, D. A. Trujillo, and K. M. Havstad. 2004. Land management in the American Southwest: a state-and-transition approach to ecosystem complexity. Environmental Management 34(1):38-51.
Callaway, R. M., and J. M. Vivanco. 2007. Invasion of plants into native plant communities using the underground information superhighway. Allelopathy Journal 19:143-151.
DiTomaso, J. M. 2000. Invasive weeds in rangelands: Species, impacts, and management. Weed Science 48:255-265.
Herrick, J. E., J. W. Van Zee, K. M. Havstad, L. M. Burkett, and W. G. Whitford. 2005. Monitoring manual for grassland, shrubland and savanna Ecosystems. Volume I Quick Start. USDA - ARS Jornada Experimental Range, Las Cruces, New Mexico.
Herrick, J. E., J. W. Van Zee, K. M. Havstad, L. M. Burkett, and W. G. Whitford. 2005. Monitoring manual for grassland, shrubland and savanna Ecosystems. Volume II: Design, supplementary methods and interpretation. USDA - ARS Jornada Experimental Range, Las Cruces, New Mexico.
Hobbs, R. J., and S. E. Humphries. 1995. An integrated approach to the ecology and management of plant invasions. Conservation Biology 9:761-770.
Lacey, J. R., C. B. Marlow, and J. R. Lane. 1989. Influence of spotted knapweed (Centaurea maculosa) on surface runoff and sediment yield. Weed Technology 3:627-631.
Launchbaugh, K. L., R. J. Daines, and J. W. Walker. [Eds.] 2006. Targeted grazing: a natural approach to vegetation management and landscape enhancement. Centennial, CO, USA: American Sheep Industry Association (available online at www.cnr.uidaho.edu/rx-grazing/Handbook.htm)
Lesica, P., and Cooper, S. V. 1997. Presettlement vegetation of southern Beaverhead County, Montana. Unpublished report to the State Office, Bureau of Land Management, and Beaverhead-Deerlodge National Forest. Montana Natural Heritage Program, Helena, MT. 35 pp.
Lowe, P. N., W. K. Laurenroth, and I. C. Burke. 2003. Effects of nitrogen availability on competition between Bromus tectorum and Bouteloua gracilis. Plant Ecology 167:247-254.
Mack, R. N., D. Simberloff, W. M. Lonsdale, H. Evans, M. Clout, and F. A. Bazzaz. 2000. Biotic invasions: causes, epidemiology, global consequences, bunchgrass ranges in southern Idaho. Journal of Range Management 24:407-410.
Masters, R. A., and R. L. Sheley. 2001. Principles and practices for managing rangeland invasive plants. Journal of Range Management 54: 502-517.
Norton, J. B., T. A. Monaco, J. M. Norton, D. A. Johnson, and T. A. Jones. 2004. Soil morphology and organic matter dynamics under cheatgrass and sagebrush-steppe plant communities. Journal of Arid Environments 57:445-466.
NRCS. 2008. National Range and Pasture Handbook. Chapter 3, Section 1, Montana Supplement: Montana Rangeland Ecological Site Key – Version 8.2.
NRCS. 2008. (electronic) Field Office Technical Guide. Available online at http://efotg.nrcs.usda.gov/efotg_locator.aspx?map=MT
NRCS. 2009. Plant Guide: Cheatgrass. Prepared by Skinner et al., National Plant Data Center.
Ogle, S., W. Reiners, and K. Gerow. 2003. Impacts of exotic annual brome grasses (Bromus spp.) on ecosystem properties of northern mixed grass prairie. Am. Midl. Nat 149:46-58.
Pellant, M. 1996. Cheatgrass: The invader of the West. Bureau of Land Management, Idaho State Office, 22 pp.
Pellant, M., P. Shaver, D. A. Pyke, and J. E. Herrick. 2005. Interpreting indicators of rangeland health. Version 4. Technical Reference 1734-6. USDI-BLM.
Pimentel, D., L. Lach, R. Zuniga, and D. Morrison. 2000. Environmental and economic costs of nonindigenous species in the United States. Bioscience 50:53-65.
Pokorny, M. L., R. L. Sheley, C. A. Zabinski, R. E. Engel, A. J. Svejcar, and J. J. Borkowski. 2005. Plant functional group diversity as a mechanism for invasion resistance. Restoration Ecology 13(3): 1-12.
Ricketts, M. J., R. S. Noggles, and B. Landgraf-Gibbons. 2004. Pryor Mountain Wild Horse Range Survey and Assessment. USDA-Natural Resources Conservation Service.
Ross, R. L., E. P. Murray, and J. G. Haigh. 1973. Soil and vegetation of near-pristine sites in Montana. USDA Soil Conservation Service, Bozeman, MT
Schoeneberger, P. J., D. A. Wysocki, E. C. Benham, and W. D. Broderson. [Edss.] 2002. Field book for describing and sampling soils, Version 2.0. Natural Resources Conservation Service, National Soil Survey Center, Lincoln, NE. (http://soils.usda.gov/technical/fieldbook/)
Sheley, R. L., B. E. Olson, and C. Hoopes. 2005. Impacts of noxious weeds. Pulling together against weeds. Published by Montana’s Statewide Noxious Weed Awareness and Education Program.
Stringham, T. K. and W. C. Krueger. 2001. States, transitions, and thresholds: Further refinement for rangeland applications. Agricultural Experiment Station, Oregon State University. Special Report 1024.
Stringham, T. K., W. C. Kreuger, and P. L Shaver. 2003. State and transition modeling: an ecological process approach. Journal of Range Management 56(2):106-113.USDA, NRCS. 1997. National Range and Pasture Handbook. (http://www.glti.nrcs.usda.gov/technical/publications/nrph.html)
U.S. Department of Agriculture, Natural Resources Conservation Service (USDA/NRCS). 2007. The PLANTS Database (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.
USDA/NRCS Soil survey manuals for appropriate counties within MLRA 44A.
Walker, L. R. and S. D. Smith. 1997. Impacts of invasive plants on community and ecosystem properties. p. 69-86. In: J. O. Luken, and J. W. Thieret. [Eds.] Assessment and management of plant invasions. Springer, New York, N.Y.
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Approval
Kirt Walstad, 3/08/2022
Rangeland health reference sheet
Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.
Author(s)/participant(s) W. Lujan, N. Svendsen, J. Alexander, K. Walstad, J. Siddoway, M. Hansen Contact for lead author NRCS Missoula Area Office Date 03/01/2010 Approved by Approval date Composition (Indicators 10 and 12) based on Annual Production Indicators
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Number and extent of rills:
Slopes on this site are typically less than 10%, but may range as high as 55%. Rills should not occur in the Taller Bunchgrass State on slopes less than 15%. As slope increases so will the number and length of rills. Rills on steep slopes (>35%) should be <5 feet long. -
Presence of water flow patterns:
Water flow patterns are generally not evident in the reference state especially on slopes <4%. Following occasional (5 – 30 % probability), heavy thunderstorms and winter thaw events, short, sinuous, discontinuous flow patterns may be apparent on 8% -35% slopes. On steeper slopes (>35%) water flow patterns may become more evident and there may be areas which show accumulations of litter due to water movement. -
Number and height of erosional pedestals or terracettes:
None to very slight. Occasionally pedestals up to 0.5 inches may be encountered. As slopes increase pedestals may become more evident. -
Bare ground from Ecological Site Description or other studies (rock, litter, lichen, moss, plant canopy are not bare ground):
Bare ground should not exceed 10% - bare areas tend to be inconspicuous and not connected. -
Number of gullies and erosion associated with gullies:
Gullies should not occur in the Taller Bunchgrass State. If there is evidence of past erosion that has created gullies, these areas should be stabilized and have no active erosion. -
Extent of wind scoured, blowouts and/or depositional areas:
Appearance or evidence of these erosional features on the landscape would not be present on this site. -
Amount of litter movement (describe size and distance expected to travel):
Litter will be evident across this site representing organic debris from the vegetation of the functional/structural groups and will not move. A severe convection storm or a significant thaw event could cause litter to move short distances, especially on slopes greater than 6%. -
Soil surface (top few mm) resistance to erosion (stability values are averages - most sites will show a range of values):
Resistance to erosion will be high with soil stability values of 5 or 6; areas of bare soil on this site may have values between 3 and 5 if not under plant canopy. -
Soil surface structure and SOM content (include type of structure and A-horizon color and thickness):
Structure is granular at the soil surface. Organic matter is about 1.5%. The surface horizon is 4 to 8 inches thick. -
Effect of community phase composition (relative proportion of different functional groups) and spatial distribution on infiltration and runoff:
The reference plant community (1.1) is dominated by rough fescue which will maximize infiltration and minimize runoff throughout the site. With the increase of Idaho fescue in Plant community (1.2) infiltration may slightly decrease and runoff may slightly increase but overall this plant community will have only minor affects on infiltration and runoff. -
Presence and thickness of compaction layer (usually none; describe soil profile features which may be mistaken for compaction on this site):
A compaction layer would not be expected on this ecological site. -
Functional/Structural Groups (list in order of descending dominance by above-ground annual-production or live foliar cover using symbols: >>, >, = to indicate much greater than, greater than, and equal to):
Dominant:
Plant community 1.1 - Taller cool season bunchgrasses (Idaho fescue) >> mid-stature cool season bunchgrasses (Idaho fescue) > cool season rhizomatous grasses (western wheatgrass), shortgrasses (prairie junegrass) and grasslikes (sedges) = perennial forbs > shrubs. Plant community 1.2 – Rough fescue and Idaho fescue share dominance – the other functional/structural groups will remain the same in descending order.Sub-dominant:
Other:
Additional:
-
Amount of plant mortality and decadence (include which functional groups are expected to show mortality or decadence):
Plant mortality for all functional groups will be low, but there will be some natural mortality of functional groups over time. Prolonged droughts and/or excessive rest may show increases in mortality and decadence for all plant groups. -
Average percent litter cover (%) and depth ( in):
Note: the majority of the litter in the plant community in the Taller Bunchgrass State will be non-persistent. -
Expected annual annual-production (this is TOTAL above-ground annual-production, not just forage annual-production):
1100 #/acre – 1700 #/acre for the reference community (1.1) with a RV of 1350 #/acre. Production varies based on effective precipitation and natural variability of soil properties for this ecological site. -
Potential invasive (including noxious) species (native and non-native). List species which BOTH characterize degraded states and have the potential to become a dominant or co-dominant species on the ecological site if their future establishment and growth is not actively controlled by management interventions. Species that become dominant for only one to several years (e.g., short-term response to drought or wildfire) are not invasive plants. Note that unlike other indicators, we are describing what is NOT expected in the reference state for the ecological site:
Cheatgrass, knapweed spp., leafy spurge, sulphur cinquefoil, dalmatian toadflax, houndstongue, whitetop, Canada thistle, Japanese brome, broom snakeweed, fringed sagewort, salsify and dandelion. -
Perennial plant reproductive capability:
All native plants are capable of reproducing sexually and/or vegetatively.
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