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Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.
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Figure 1. Mapped extent
Areas shown in blue indicate the maximum mapped extent of this ecological site. Other ecological sites likely occur within the highlighted areas. It is also possible for this ecological site to occur outside of highlighted areas if detailed soil survey has not been completed or recently updated.
MLRA notes
Major Land Resource Area (MLRA): 040X–Sonoran Basin and Range
AZ 40.1 – Upper Sonoran Desert
Elevations range from 2000 to 3200 feet and precipitation averages 10 to 13 inches per year. Vegetation includes saguaro, palo verde, mesquite, creosotebush, triangle bursage, prickly pear, cholla, limberbush, wolfberry, bush muhly, threeawns, ocotillo, and globe mallow. The soil temperature regime is thermic and the soil moisture regime is typic aridic. This unit occurs within the Basin and Range Physiographic Province and is characterized by numerous mountain ranges that rise abruptly from broad, plain-like valleys and basins. Igneous and metamorphic rock classes dominate the mountain ranges and sediments filling the basins represent combinations of fluvial, lacustrine, colluvial and alluvial deposits.Associated sites
R040XA110AZ Limy Slopes 10"-13" p.z.
R040XA111AZ Limy Upland 10"-13" p.z.
R040XA127AZ Gypsum Slopes 10"-13" p.z.
Similar sites
R040XA108AZ Limy Fan 10"-13" p.z.
R041XB219AZ Gypsum Upland 8-12" p.z.
Table 1. Dominant plant species
Tree (1) Parkinsonia microphylla
Shrub (1) Larrea tridentata
(2) Acacia constrictaHerbaceous (1) Aristida purpurea
(2) Cevallia sinuataPhysiographic features
This site occurs in the upper elevations of the Sonoran Desert province in southern Arizona. It occurs on fan terraces formed on relict lake bed sediments.
Table 2. Representative physiographic features
Landforms (1) Alluvial flat
(2) Fan remnant
Flooding frequency None Ponding frequency None Elevation 2600 – 3600 ft Slope 1 – 5 % Aspect Aspect is not a significant factor Climatic features
Precipitation in the sub resource area ranges from 10 to 13 inches in the southern part, along the Mexican border with elevations from about 1900 to 3200 feet. Precipitation in the northern part of the resource area ranges from 11 to 14 inches with elevations from about 1700 to 3500 feet. Winter-summer rainfall ratios range from 40%-60% in the southern portions of the land resource unit, to 50%-50% in the central portions, to 60%-40% in the northern part of the land resource unit. As one moves from east to west in this resource area rains become slightly more unpredictable and variable with Coefficients of Variation of annual rainfall equal to 29% at Tucson and 36% at Carefree. Summer rains fall July through Sept., originate in the Gulf of Mexico, and are convective, usually brief, intense thunderstorms. Cool season moisture tends to be frontal, originating in the Pacific and Gulf of California. This winter precipitation falls in widespread storms with long duration and low intensity. Snow is rare and seldom lasts more than an hour or two. May and June are the driest months of the year. Humidity is generally very low.
Winter temperatures are mild, with very few days recording freezing temperatures in the morning. Summer temperatures are warm to hot, with several days in June and July exceeding 105 degrees F.
Both the spring and the summer growing seasons are equally important for perennial grass, forb and shrub growth. Cool and warm season annual forbs and grasses can be common in their respective seasons with above average rainfall. Perennial forage species can remain green throughout the year with available moisture.Table 3 Representative climatic features
Frost-free period (average) 270 days Freeze-free period (average) 0 days Precipitation total (average) 10 in BarLineFigure 2. Monthly precipitation range
BarLineFigure 3. Monthly average minimum and maximum temperature
">Influencing water features
There are no water features associated with this site.
Soil features
These soils are well drained, variable in texture, stratified and high in soluble gypsum (5-30% by volume). They are moderately deep to deep and underlain by gypsum deposits in places. Coarse fragments in the soil profile are usually large gypsum crystals. These soils have formed in relict lacustrine deposits and may be slightly saline and sodic as well. They are not flooded. Soil series mapped on this site include: in SSA-666 Northwest Cochise County MU's 247-Contention & calcigypsids.
Table 4. Representative soil features
Surface texture (1) Silt loam
(2) Fine sandy loam
(3) Clay
Family particle size (1) Loamy
Drainage class Well drained Permeability class Moderate to slow Soil depth 60 in Surface fragment cover <=3" 0 – 20 % Surface fragment cover >3" 0 – 5 % Available water capacity
(0-40in)4.2 – 6.6 in Calcium carbonate equivalent
(0-40in)0 – 20 % Electrical conductivity
(0-40in)0 – 2 mmhos/cm Sodium adsorption ratio
(0-40in)0 – 2 Soil reaction (1:1 water)
(0-40in)7.4 – 8.4 Subsurface fragment volume <=3"
(Depth not specified)0 – 20 % Subsurface fragment volume >3"
(Depth not specified)0 – 5 % Ecological dynamics
The plant communities found on an ecological site are naturally variable. Composition and production will vary with yearly conditions, location, aspect and the natural variability of the soils. The Historical Climax Plant Community represents the natural potential plant community found on relict or relatively undisturbed areas of this site. Other plant communities described here represent plant communities that are known to occur when the site is disturbed by factors such as fire, grazing and drought.
Production data provided in this site description is standardized to air dry weight at the end of the summer growing season. The plant communities described in this site description are based on near normal rainfall years.
NRCS uses a Similarity Index to compare existing plant communities to the plant communities described here. Similarity index is determined by comparing the production and composition of a plant community to the production and composition described in the site description. To determine similarity index, compare the production (air dry weight) of each species to that shown in the plant community description. For each species, count no more than the maximum amount shown for the species, and for each group, count no more than the amount shown for that group. Divide the resulting total by the total, normal year, production shown in the plant community description. If the rainfall has been significantly above or below normal, use the total production shown for above or below normal years. If the field data is not collected at the end of the summer growing season, then the field data must be corrected to the end of year production before comparing it to the site description. The growth curve can be used as a guide for estimating production at the end of the summer growing season.
The historic native state includes the native plant communities that occur on the site, including the historic climax plant community. This state includes other plant communities that naturally occupy the site following fire, drought, flooding, herbivores and other natural disturbances. The historic climax plant community represents the natural climax community that eventually reoccupies the site with proper management and a return to near normal conditions and /or equilibrium.
State and transition model
Custom diagramStandard diagram
Figure 4. Gypsum upland 10-13" pz
More interactive model formats are also available. View Interactive Models
More interactive model formats are also available. View Interactive Models
Click on state and transition labels to scroll to the respective textEcosystem states
State 1 submodel, plant communities
State 2 submodel, plant communities
State 3 submodel, plant communities
State 4 submodel, plant communities
State 1
Historic Climax Plant CommunityCommunity 1.1
Historic Climax Plant CommunityThis plant community is dominated by creosote bush and whitethorn acacia with lesser amounts of trees like mesquite and paloverde. Annual grasses and forbs are an important part of the plant community. Cryptogams are very common on this site, often completely colonizing outcrops of gypsum. The site is extremely susceptible to soil piping and sheet, rill and gully erosion due to high concentrations of soluble gypsum in the surface soil.
Figure 5. Annual production by plant type (representative values) or group (midpoint values)
Table 5. Annual production by plant type
Plant type Low
(lb/acre)Representative value
(lb/acre)High
(lb/acre)Shrub/Vine 100 250 340 Grass/Grasslike 2 50 165 Forb 2 50 160 Tree 0 20 50 Total 104 370 715 Table 6. Soil surface cover
Tree basal cover 0-0% Shrub/vine/liana basal cover 0% Grass/grasslike basal cover 0-0% Forb basal cover 0-0% Non-vascular plants 0% Biological crusts 30-80% Litter 10-30% Surface fragments >0.25" and <=3" 10-30% Surface fragments >3" 0-10% Bedrock 0-10% Water 0% Bare ground 20-70% Table 7. Canopy structure (% cover)
Height Above Ground (ft) Tree Shrub/Vine Grass/
GrasslikeForb <0.5 – 0-1% 0-5% 1-10% >0.5 <= 1 – 0-2% 0-5% 0-5% >1 <= 2 – 1-2% 0-5% 0-1% >2 <= 4.5 – 5-10% 0-2% 0-1% >4.5 <= 13 0-2% 0-5% – – >13 <= 40 – – – – >40 <= 80 – – – – >80 <= 120 – – – – >120 – – – – Figure 6. Plant community growth curve (percent production by month). AZ4013 , 40.1 10-13" p.z. other sites. Growth begins in the late winter, goes semi-dormant in the drought period of late May through early July, growth continues in the summer through early fall..
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec J F M A M J J A S O N D 0 5 10 20 15 0 5 20 15 5 5 0 State 2
Shrubs, cacti and fireCommunity 2.1
Shrubs, cacti and fireThis state occurs where the native tree, cacti and shrub cover has been impacted by a single hot season fire. This will occur only in El Nino years when annual forbs and grasses make enough fuel for a fire. Areas of this site will not burn completely due to patches of soil dominated by gypsum outcrop and high cryptogam covers which will not have fine fuel.
The fire impacts paloverde, saguaro and other cacti species. Shrubs like creosote and whitethorn may sprout and recover from the fire.State 3
Exotic annualsCommunity 3.1
Exotic annualsThis state occurs where non-native annuals have invaded the native plant community. Species like red brome, schismus and native annuals dominate the understory of the plant community. Areas of the site adjacent urban areas and near heavily travelled roads may burn more often due to increased chance of ignition.
State 4
ErodedCommunity 4.1
ErodedThis state occurs where the site has been graded, bladed or mined and the soil surface (gravelly sandyloam) has been disturbed. The subsurface is extremely erodible due to high concentrations of souble gypsum in the soil profile. Vehicle and ATV traffic may contribute to rilling and gully formation.
Additional community tables
Table 8. Community 1.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Grass/Grasslike1 Dominant perennial grasses 1–50 bush muhly MUPO2 Muhlenbergia porteri 1–15 – whiplash pappusgrass PAVA2 Pappophorum vaginatum 0–10 – purple threeawn ARPU9 Aristida purpurea 0–10 – Parish's threeawn ARPUP5 Aristida purpurea var. parishii 0–10 – false Rhodes grass TRCR9 Trichloris crinita 0–10 – alkali sacaton SPAI Sporobolus airoides 0–5 – spike dropseed SPCO4 Sporobolus contractus 0–5 – big sacaton SPWR2 Sporobolus wrightii 0–5 – 2 Misc. perennial grasses 1–15 spidergrass ARTE3 Aristida ternipes 0–5 – spidergrass ARTEG Aristida ternipes var. gentilis 0–5 – low woollygrass DAPU7 Dasyochloa pulchella 0–5 – nineawn pappusgrass ENDE Enneapogon desvauxii 0–5 – sand dropseed SPCR Sporobolus cryptandrus 0–2 – mesa dropseed SPFL2 Sporobolus flexuosus 0–2 – tobosagrass PLMU3 Pleuraphis mutica 0–1 – plains bristlegrass SEVU2 Setaria vulpiseta 0–1 – Arizona cottontop DICA8 Digitaria californica 0–1 – Havard's threeawn ARHA3 Aristida havardii 0–1 – blue threeawn ARPUN Aristida purpurea var. nealleyi 0–1 – 3 Annual grasses 0–100 sixweeks threeawn ARAD Aristida adscensionis 0–50 – needle grama BOAR Bouteloua aristidoides 0–50 – sixweeks grama BOBA2 Bouteloua barbata 0–50 – mucronate sprangeltop LEPAB Leptochloa panicea ssp. brachiata 0–20 – prairie threeawn AROL Aristida oligantha 0–15 – Rothrock's grama BORO2 Bouteloua rothrockii 0–10 – sixweeks fescue VUOC Vulpia octoflora 0–10 – Mexican panicgrass PAHI5 Panicum hirticaule 0–5 – Bigelow's bluegrass POBI Poa bigelovii 0–2 – Arizona signalgrass URAR Urochloa arizonica 0–2 – canyon cupgrass ERLE7 Eriochloa lemmonii 0–2 – desert lovegrass ERPEM Eragrostis pectinacea var. miserrima 0–2 – tufted lovegrass ERPEP2 Eragrostis pectinacea var. pectinacea 0–2 – Mexican sprangletop LEFUU Leptochloa fusca ssp. uninervia 0–1 – delicate muhly MUFR Muhlenbergia fragilis 0–1 – littleseed muhly MUMI Muhlenbergia microsperma 0–1 – Arizona brome BRAR4 Bromus arizonicus 0–1 – Forb4 Perennial forbs 1–10 stinging serpent CESI Cevallia sinuata 1–5 – desert trumpet ERIN4 Eriogonum inflatum 0–5 – brownplume wirelettuce STPA4 Stephanomeria pauciflora 0–5 – desert globemallow SPAM2 Sphaeralcea ambigua 0–2 – weakleaf bur ragweed AMCO3 Ambrosia confertiflora 0–2 – tuber anemone ANTU Anemone tuberosa 0–1 – dense ayenia AYMI Ayenia microphylla 0–1 – hairyseed bahia BAAB Bahia absinthifolia 0–1 – desert marigold BAMU Baileya multiradiata 0–1 – whitemargin sandmat CHAL11 Chamaesyce albomarginata 0–1 – leatherweed CRPOP Croton pottsii var. pottsii 0–1 – dwarf desertpeony ACNA2 Acourtia nana 0–1 – brownfoot ACWR5 Acourtia wrightii 0–1 – trailing windmills ALIN Allionia incarnata 0–1 – San Pedro daisy LAPO4 Lasianthaea podocephala 0–1 – Parry's false prairie-clover MAPA7 Marina parryi 0–1 – lacy tansyaster MAPIP4 Machaeranthera pinnatifida ssp. pinnatifida var. pinnatifida 0–1 – wishbone-bush MILAV Mirabilis laevis var. villosa 0–1 – desert tobacco NIOB Nicotiana obtusifolia 0–1 – Coues' cassia SECO10 Senna covesii 0–1 – silverleaf nightshade SOEL Solanum elaeagnifolium 0–1 – pricklyleaf dogweed THAC Thymophylla acerosa 0–1 – rue of the mountains THTE2 Thamnosma texana 0–1 – 5 Annual forbs 1–150 California poppy ESCAM Eschscholzia californica ssp. mexicana 0–25 – Gordon's bladderpod LEGO Lesquerella gordonii 0–25 – western tansymustard DEPI Descurainia pinnata 0–20 – Coulter's lupine LUSP2 Lupinus sparsiflorus 0–20 – bristly fiddleneck AMTE3 Amsinckia tessellata 0–15 – thelypody THELY Thelypodium 0–15 – exserted Indian paintbrush CAEXE Castilleja exserta ssp. exserta 0–10 – combseed PECTO Pectocarya 0–10 – manybristle chinchweed PEPA2 Pectis papposa 0–10 – desert Indianwheat PLOV Plantago ovata 0–10 – flatcrown buckwheat ERDE6 Eriogonum deflexum 0–10 – miniature woollystar ERDI2 Eriastrum diffusum 0–10 – cryptantha CRYPT Cryptantha 0–5 – shaggyfruit pepperweed LELA Lepidium lasiocarpum 0–5 – intermediate pepperweed LEVIM Lepidium virginicum var. medium 0–5 – coastal bird's-foot trefoil LOSAB Lotus salsuginosus var. brevivexillus 0–5 – slender goldenweed MAGR10 Machaeranthera gracilis 0–5 – Nuttall's povertyweed MONU Monolepis nuttalliana 0–5 – wheelscale saltbush ATEL Atriplex elegans 0–5 – Coulter's spiderling BOCO2 Boerhavia coulteri 0–5 – carelessweed AMPA Amaranthus palmeri 0–5 – lyreleaf jewelflower STCAA Streptanthus carinatus ssp. arizonicus 0–5 – woollyhead neststraw STMI2 Stylocline micropoides 0–2 – woolly tidestromia TILA2 Tidestromia lanuginosa 0–2 – fringed redmaids CACI2 Calandrinia ciliata 0–2 – white tackstem CAWR Calycoseris wrightii 0–2 – brittle spineflower CHBR Chorizanthe brevicornu 0–2 – hyssopleaf sandmat CHHY3 Chamaesyce hyssopifolia 0–2 – Esteve's pincushion CHST Chaenactis stevioides 0–2 – green carpetweed MOVE Mollugo verticillata 0–2 – tanseyleaf tansyaster MATA2 Machaeranthera tanacetifolia 0–2 – desert evening primrose OEPR Oenothera primiveris 0–2 – Florida pellitory PAFL3 Parietaria floridana 0–2 – phacelia PHACE Phacelia 0–2 – hairy desertsunflower GECA2 Geraea canescens 0–2 – star gilia GIST Gilia stellata 0–2 – Arizona poppy KAGR Kallstroemia grandiflora 0–2 – hairy prairie clover DAMO Dalea mollis 0–2 – American wild carrot DAPU3 Daucus pusillus 0–2 – sorrel buckwheat ERPO4 Eriogonum polycladon 0–2 – Texas stork's bill ERTE13 Erodium texanum 0–2 – common woolly sunflower ERLA6 Eriophyllum lanatum 0–1 – Mexican fireplant EUHE4 Euphorbia heterophylla 0–1 – Arizona popcornflower PLAR Plagiobothrys arizonicus 0–1 – doubleclaw PRPA2 Proboscidea parviflora 0–1 – New Mexico plumeseed RANE Rafinesquia neomexicana 0–1 – sleepy silene SIAN2 Silene antirrhina 0–1 – Coulter's globemallow SPCO2 Sphaeralcea coulteri 0–1 – bristly nama NAHI Nama hispidum 0–1 – glandular threadplant NEGL Nemacladus glanduliferus 0–1 – yellow tackstem CAPA7 Calycoseris parryi 0–1 – hoary bowlesia BOIN3 Bowlesia incana 0–1 – milkvetch ASTRA Astragalus 0–1 – Shrub/Vine6 Dominant shrubs 100–300 creosote bush LATR2 Larrea tridentata 50–200 – whitethorn acacia ACCO2 Acacia constricta 0–100 – ocotillo FOSP2 Fouquieria splendens 0–15 – 7 Miscellaneous shrubs 0–15 ocotillo FOSP2 Fouquieria splendens 0–15 – fourwing saltbush ATCA2 Atriplex canescens 0–5 – longleaf jointfir EPTR Ephedra trifurca 0–1 – American tarwort FLCE Flourensia cernua 0–1 – cattle saltbush ATPO Atriplex polycarpa 0–1 – Warnock's snakewood COWA Condalia warnockii 0–1 – catclaw acacia ACGR Acacia greggii 0–1 – Wright's beebrush ALWR Aloysia wrightii 0–1 – crown of thorns KOSP Koeberlinia spinosa 0–1 – water jacket LYAN Lycium andersonii 0–1 – Berlandier's wolfberry LYBE Lycium berlandieri 0–1 – Arizona desert-thorn LYEX Lycium exsertum 0–1 – soaptree yucca YUEL Yucca elata 0–1 – lotebush ZIOB Ziziphus obtusifolia 0–1 – 8 Half shrubs 0–15 whitestem paperflower PSCO2 Psilostrophe cooperi 0–5 – broom snakeweed GUSA2 Gutierrezia sarothrae 0–5 – Parish's goldeneye VIPA14 Viguiera parishii 0–5 – desert zinnia ZIAC Zinnia acerosa 0–2 – pricklyleaf dogweed THAC Thymophylla acerosa 0–2 – littleleaf ratany KRER Krameria erecta 0–1 – winterfat KRLA2 Krascheninnikovia lanata 0–1 – mariola PAIN2 Parthenium incanum 0–1 – rayless goldenhead ACSP Acamptopappus sphaerocephalus 0–1 – triangle bur ragweed AMDE4 Ambrosia deltoidea 0–1 – burrobush AMDU2 Ambrosia dumosa 0–1 – brittlebush ENFA Encelia farinosa 0–1 – threadleaf snakeweed GUMI Gutierrezia microcephala 0–1 – 9 Succulents 1–20 devil's cholla GRKU Grusonia kunzei 0–10 – cactus apple OPEN3 Opuntia engelmannii 0–5 – jumping cholla CYFU10 Cylindropuntia fulgida 0–5 – Christmas cactus CYLE8 Cylindropuntia leptocaulis 1–5 – staghorn cholla CYVE3 Cylindropuntia versicolor 0–5 – tulip pricklypear OPPH Opuntia phaeacantha 0–2 – nightblooming cereus PEGR3 Peniocereus greggii 0–1 – walkingstick cactus CYSP8 Cylindropuntia spinosior 0–1 – Engelmann's hedgehog cactus ECEN Echinocereus engelmannii 0–1 – pinkflower hedgehog cactus ECFA Echinocereus fasciculatus 0–1 – candy barrelcactus FEWI Ferocactus wislizeni 0–1 – saguaro CAGI10 Carnegiea gigantea 0–1 – buck-horn cholla CYAC8 Cylindropuntia acanthocarpa 0–1 – Tree10 Trees 0–50 velvet mesquite PRVE Prosopis velutina 0–50 – yellow paloverde PAMI5 Parkinsonia microphylla 0–50 – western honey mesquite PRGLT Prosopis glandulosa var. torreyana 0–10 – Table 9. Community 2.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 10. Community 3.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Table 11. Community 4.1 plant community composition
Group Common name Symbol Scientific name Annual production () Foliar cover (%) Interpretations
Animal community
This site offers little in the way of livestock forage. It does produce limited forage of annual grasses and forbs in wet winters. High amounts of soluble gypsum in runoff water that is collected in stock ponds may have a laxative effect on livestock.
Wildlife on the site is limited to small mammals and birds and their associated predators.Hydrological functions
Medium to heavy textured soils make this a moderate producer of runoff. Soluble gypsum can lead to piping and rilling with heavy rainfall.
Recreational uses
Hunting, horseback riding, hiking, four wheeling.
Wood products
Some mesquite, catclaw and paloverde for camp and branding fires.
Other products
Gypsum is mined in many areas for material used in making sheet rock.
Supporting information
Type locality
Location 1: Pinal County, AZ Township/Range/Section T5S R17E S15 General legal description On the old Mary Cluff Ranch on the Mineral Strip, San Carlos Apache Nation. Location 2: Pinal County, AZ Township/Range/Section T6S R16E S27 General legal description On the old PZ Ranch, now Becky Stambaugh ranch. Contributors
Dan Robinett
Larry D. EllicottRangeland health reference sheet
Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.
Author(s)/participant(s) Contact for lead author Date Approved by Approval date Composition (Indicators 10 and 12) based on Annual Production Indicators
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Number and extent of rills:
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Presence of water flow patterns:
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Number and height of erosional pedestals or terracettes:
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Bare ground from Ecological Site Description or other studies (rock, litter, lichen, moss, plant canopy are not bare ground):
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Number of gullies and erosion associated with gullies:
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Extent of wind scoured, blowouts and/or depositional areas:
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Amount of litter movement (describe size and distance expected to travel):
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Soil surface (top few mm) resistance to erosion (stability values are averages - most sites will show a range of values):
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Soil surface structure and SOM content (include type of structure and A-horizon color and thickness):
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Effect of community phase composition (relative proportion of different functional groups) and spatial distribution on infiltration and runoff:
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Presence and thickness of compaction layer (usually none; describe soil profile features which may be mistaken for compaction on this site):
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Functional/Structural Groups (list in order of descending dominance by above-ground annual-production or live foliar cover using symbols: >>, >, = to indicate much greater than, greater than, and equal to):
Dominant:
Sub-dominant:
Other:
Additional:
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Amount of plant mortality and decadence (include which functional groups are expected to show mortality or decadence):
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Average percent litter cover (%) and depth ( in):
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Expected annual annual-production (this is TOTAL above-ground annual-production, not just forage annual-production):
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Potential invasive (including noxious) species (native and non-native). List species which BOTH characterize degraded states and have the potential to become a dominant or co-dominant species on the ecological site if their future establishment and growth is not actively controlled by management interventions. Species that become dominant for only one to several years (e.g., short-term response to drought or wildfire) are not invasive plants. Note that unlike other indicators, we are describing what is NOT expected in the reference state for the ecological site:
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Perennial plant reproductive capability:
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