Ecological dynamics
The historic climax plant community (HCPC) for a site in North America is the plant community that existed at the time of European immigration and settlement. It is the plant community that was best adapted to the unique combination of environmental factors associated with the site at that time. The HCPC was in dynamic equilibrium with its environment and was able to avoid displacement by the suite of disturbances and disturbance patterns (magnitude and frequency) that naturally occurred within the area occupied by the site. Natural disturbances, such as drought, fire, grazing of native fauna, and insects, were inherent in the development and maintenance of the plant community. The effects of these disturbances are part of the range of characteristics of the site that contribute to the dynamic equilibrium. Fluctuations in the plant community’s structure and function caused by the effects of these natural disturbances establish the boundaries of dynamic equilibrium. They are accounted for as part of the range of characteristics for the ecological site. The HCPC is not a precise assemblage of species for which the proportions are the same from place to place or from year to year. Variability is apparent in productivity and occurrence of individual species.
The HCPC for this ecological site has been estimated by sampling relict or relatively undisturbed sites and/or reviewing historic records.
A plant community that is subjected to abnormal disturbances and physical site deterioration or that is protected from natural influences, such as fire and grazing, for long periods seldom typifies the HCPC. Any physical site deterioration caused by the abnormal disturbance may result in the crossing of a threshold or irreversible boundary to another state, or equilibrium, for the ecological site. There may be multiple thresholds and states possible for an ecological site, determined by the type and or severity of abnormal disturbance. The known states and transition pathways for this ecological site are described in the accompanying state and transition model.
The “Plant Community Plant Species Composition” table provides a list of species and each species or group of species’ annual production in pounds per acre (air-dry weight) expected in a normal rainfall year. Low and high production yields represent the modal range of variability for that species or group of species across the extent of the ecological site.
The “Annual Production by Plant Type” table provides the median air-dry production and the fluctuations to be expected during favorable, normal, and unfavorable years.
The present plant community on an ecological site can be compared to the various common vegetation states that can exist on the site. The degree of similarity is expressed through a similarity index. To determine the similarity index, compare the production of each species to that shown in the plant community description. For each species, count no more than the maximum amount shown for the species, and for each group, count no more than the maximum shown for the group. Divide the resulting total by the total representative value shown in the “Annual Production by Plant Type” table for the reference plant community. Variations in production due to above or below normal rainfall, incomplete growing season or utilization must be corrected before comparing it to the site description. The “Worksheet for Determining Similarity Index” is useful in making these corrections. The accompanying growth curve can be used as a guide for estimating percent of growth completed.
State and transition model
Custom diagram
Standard diagram
Additional community tables
